Amphicyon longiramus 01

Ventral view of skull, UF 162723

Amphicyon longiramus

Quick Facts

Common Name: White’s bear-dog

The Florida Museum of Natural History has over 550 specimens of this species in its collection, and has the only known mounted skeleton of the species on public display.

Amphicyon longiramus was the top carnivore in Florida 16 to 18 million years ago.

Some species of bear-dogs evolved wolf-like qualities such as longer legs and tearing-teeth, while others (including White's bear-dog) evolved more bear-like qualities such as a broader body and slower gait, and crushing molars.

Its closest modern ecologic equivalent is the grizzly bear.

Age Range

Scientific Name and Classification

Amphicyon longiramus White, 1942

Source of Species Name: (longiramus is Latin for “long branch”). When Amphicyon longiramus was originally named by Dr. Theodore E. White of the Harvard Museum of Comparative Zoology, he believed that it coexisted along with another, smaller species of Amphicyon. The name was used to emphasize this species’ seemingly longer limbs and skull (many bony elements are generally referred to as a “ramus” in anatomy). The two species originally named by Dr. White have since been combined under the name Amphicyon longiramus, with more specimens making it clear that the initial variation he observed was within a single sexually dimorphic species (Olsen, 1960).

Scientific classification: Mammalia, Eutheria, Laurasiatheria, Carnivora, Caniformia, Arctoidea, Amphicyonidae, Amphicyoninae

Alternate Scientific Name: Amphicyon intermedius White, 1940 (not Amphicyon intermedius H. von Meyer, 1849)

Amphicyon longiramus, map

Figure 1. Map of Florida, highlights indicating counties where fossils of this species has been found.

Overall Geographic Range

Known primarily from the Thomas Farm Site in north-central Florida and several localities in southeastern Texas. Emry and Eshelman (1998) reported specimens of “Amphicyon intermedius” from the Pollack Farm Local Fauna of Delaware, which are similar to the same elements of Amphicyon longiramus from Florida. This occurrence indicates that Amphicyon longiramus may have been distributed throughout much of the eastern half of North America during the early Miocene.

Florida Fossil Occurrences

Florida fossil sites with Amphicyon longiramus


Amphicyon longiramus was the top carnivore of Florida’s early Miocene (Hulbert, 2001), with the largest body size among predatory animals preserved at the Thomas Farm locality. Body mass estimation among amphicyonids is notoriously difficult (Figueirido et al., 2011), due in part to the disproportionately long skulls in this group—a feature that is exaggerated in Amphicyon longiramus (Olsen, 1958); yet Amphicyon longiramus clearly dominates in size over contemporaneous carnivores. This makes identification of skeletal elements relatively easy for researchers.

Amphicyon longiramus is one of the oldest members of the subfamily Amphicyoninae in Florida. Along with the smaller amphicyonine Cynelos caroniavorus, also found at Thomas Farm (Hulbert, 2001), Amphicyon longiramus is thought to have dispersed from Europe into North America about 18 million years ago (Hunt, 1998). While other beardog species were present in Florida from the late Oligocene to the late Miocene (Hulbert, 2001), most are known only from sparse material. Amphicyon longiramus is the only species for which there is ample fossil evidence, and it is therefore the only Florida amphicyonid that has been thoroughly described (Olsen, 1958, 1960). The holotype of the species, along with other specimens, is housed in the Harvard Museum of Comparative Zoology. The Florida Museum of Natural History has over 550 specimens in its collection, and has the only known mounted skeleton of the species on public display.

Amphicyonids encompass a wide variety of carnivorous mammal ecomorphologies that are today filled by canids and ursids (Figuerido et al., 2011). Like modern carnivores, many species of bear-dogs (including Amphicyon longiramus) were highly sexually dimorphic (Hunt, 1998; Figs. 2-3), with males being much larger than females. Fossil burrows indicating denning behavior in large amphicyonids have been found in Nebraska (Hunt et al., 1983).

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Figure 2. UF 162723, skull of Amphicyon longiramus, with diagnostic auditory region highlighted, in left lateral, dorsal, and ventral views. This relatively small individual is most likely a female.
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Figure 3. Right lower canine teeth of Amphicyon longiramus from Thomas Farm demonstrating sexual dimorphism in body size. A, UF 271019, male; B, UF 154301, female.

Interestingly, two locomotive types seem to have developed within the family Amphicyonidae, emphasizing two different feeding strategies. Some amphicyonids were digitigrade, long-limbed, and had narrow limb articulations, making them effective running predators. One example is Borocyon robustum, a well-known amphicyonid in western North America (Hunt, 1998, 2009). These bear-dogs emphasized a shearing dentition excellent for slicing meat, and were the ecological equivalents of wolves. In contrast, other species of bear-dogs, such as Amphicyon longiramus, had plantigrade feet and robust limbs with broad articulations (Figs. 3-4), giving them a stance much more similar to modern bears (Hunt, 1998). These heavy, slow-gaited amphicyonines tended to have broader molars, well-designed for bone crushing, such as the upper first molar (M1) of Amphicyon longiramus pictured in Figure 6. Its closest modern ecologic equivalent is the grizzly bear.

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Figure 4. Right femur (UF 176191) of Amphicyon longiramus. A, Anterior view, with robust portions of the femur enclosed in white brackets; B, posterior view; C, lateral view; D, medial view with fovea capitis highlighted. The fovea capitis is positioned more posteriorly in amphicyonids adapted for running.

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Figure 5. Right astragalus (UF/FGS 6336) of Amphicyon longiramus. A, Medial view; B, anterior view, with broad astragalar trochlea for articulation with the tibia enclosed in white brackets. The trochlea is much narrower in amphicyonids adapted for running; C, posterior view.
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Figure 5. UF 165019, left upper M1 of Amphicyon longiramus from Thomas Farm. Above, lateral or buccal view; below, occlusal view.


  • Original Author(s): Paul E. Morse
  • Original Completion Date: October 5, 2012
  • Editor(s) Name(s): Richard C. Hulbert Jr. and Natali Valdes
  • Last Updated On: February 25, 2015
Scientific References (Click to view)


This material is based upon work supported by the National Science Foundation under Grant Number CSBR 1203222, Jonathan Bloch, Principal Investigator. Any opinions, findings, and conclusions or recommendations expressed in this material are those of the author(s) and do not necessarily reflect the views of the National Science Foundation. 

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