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Whitten, W.M., N.H. Williams, & M.W. Chase. 2000. Subtribal and generic relationships of Maxillarieae (Orchidaceae) with emphasis on Stanhopeinae: combined molecular evidence. American Journal of Botany 87: 1842-1856.

Whitten, W.M., N. H. Williams, R. L. Dressler, G. Gerlach, and F. Pupulin. 2005. Generic relationships of Zygopetalinae (Orchidaceae: Cymbidieae): combined molecular evidence. Lankesteriana 5: 87-108.

Williams, N.H. & W.M. Whitten. 2003. Molecular phylogenetics and generic concepts in Maxillarieae (Orchidaceae). Invited lecture, Primero Congreso Internacional de Orquideologia Neotropical, San Jose, Costa Rica

Williams, N.H., M. W. Chase, T. Fulcher, & W. M. Whitten. 2001. Molecular systematics of the Oncidiinae based on evidence from four DNA sequence regions: expanded circumscriptions of Cyrtochilum, Erycina, Otoglossum, and Trichocentrum and a new genus (Orchidaceae). Lindleyana 16: 113-139.

Williams, N.H., M. W. Chase, & W. M. Whitten. 2001. Phylogenetic position of Miltoniopsis, Caucaea, a new genus, Cyrtochiloides, and relationship of Oncidium phymatochilum based on nuclear and chloroplast DNA sequence data (Orchidaceae: Oncidiinae). Lindleyana 16: 95-114.

Williams, N. H. & W. M. Whitten. 2001. Checking an orchid hybrid's background: The use of molecular data in determining parentage of hybrids. Orchids 70: 1056-1061.

Williams, N.H., W.M. Whitten, and R. L. Dressler. 2005. Molecular systematics of Telipogon (Orchidaceae: Oncidiinae) and its allies: nuclear and plastid dna sequence data. Lankesteriana 5: 163-184.

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