Phylogenetics of Maxillariinae (Orchidaceae)

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We recently completed our study of the molecular systematics of Maxillaria and its relatives and published the results in the following paper

Whitten, W.M., M. A. Blanco, N. H. Williams, S. Koehler, G. Carnevali, R. B. Singer, L. Endara, & K. M. Neubig. 2007. Molecular Phylogenetics of Maxillaria and Related Genera (Orchidaceae: Cymbidieae) Based Upon Combined Molecular Data Sets. American Journal of Botany 94: 1860-1889.

ABSTRACT. Phylogenetic relationships within the orchid subtribe Maxillariinae sensu Dressler (1993) with emphasis on Maxillaria s.l. were inferred using parsimony analyses of individual and combined DNA sequence data. We analyzed nrITS DNA, the plastid matK gene and flanking trnK intron, and the plastid atpB-rbcL intergenic spacer for 619 individuals representing ca. 354 species. The plastid rpoC1 gene (ca. 2600 bp) was sequenced for 84 selected taxa and combined in a more limited analysis with the other data sets to provide greater resolution in the spine of the trees. A well-resolved and supported consensus was found, for which most clades were present in more than one individual analysis. All the currently recognized minor genera of "core" Maxillariinae (Anthosiphon, Chrysocycnis, Cryptocentrum, Cyrtidiorchis, Mormolyca, Pityphyllum, and Trigonidium) are embedded within a polyphyletic Maxillaria s.l. Our results support the recognition of a smaller, more restricted Maxillaria, of some previously published segregate genera (Brasiliorchis, Camaridium, Christensonella, Heterotaxis, Ornithidium, Sauvetrea), and of several novel clades at the generic level. These revised monophyletic generic and infrageneric concepts will hopefully minimize further nomenclatural changes, encourage monographic studies, and facilitate more focused analyses of character evolution within Maxillariinae.

We published the nomenclatorial changes in another paper

Blanco, M. A. G. Carnevali, W. M. Whitten, R. Singer, S. Koehler, N. H. Williams, I. Ojeda, K. Neubig, & L. Endara. 2007. Generic Realignments in Maxillariinae (Orchidaceae). Lankesteriana 7(3): 515-537.

You can access the pdf here Generic Realignments in Maxillariinae (Orchidaceae).

ABSTRACT. A recent phylogenetic analysis of four DNA regions for ca. 354 species of core Maxillariinae strongly indicate that the genus Maxillaria, as traditionally circumscribed, is grossly polyphyletic. We present a new phylogenetic classification for core Maxillariinae that recognizes 17 genera. Necessary realignments include: 1) resurrection of the genera Camaridium, Heterotaxis, and Ornithidium; 2) recognition of the recent segregates Brasiliorchis (=Maxillaria sect. Repentes), Christensonella (=Maxillaria sect.Urceolatae), Nitidobulbon (in press), and a recircumscribed Sauvetrea (=Maxillaria sect. Trigonae); 3) adoption of the new genera Inti (=Maxillaria sect.Polyphyllae), Mapinguari, Maxillariella (=Maxillaria sections Ebulbes and Erectae), and Rhetinantha; 4) transfers from Maxillaria sect. Reflexae to Ornithidium, and Maxillaria sect. Rufescens to Mormolyca; and 5) synonymizing of the genera Adamanthus, Pseudomaxillaria, Psittacoglossum, and Sepalosaccus (under Camaridium), Anthosiphon (under Cryptocentrum), Chrysocycnis (under Mormolyca), Dicrypta, Marsupiaria, and Pentulop (under Heterotaxis), and Laricorchis, Neo-urbania, and Siagonanthus (under Ornithidium). Some new synonyms at the specific level are also presented.

We have not updated the list below to reflect the changes in the paper above, but leave this here to show the thinking on Maxillaria and its relatives when we began our project.

The list below was compiled before any molecular work was begun on the Maxillariinae and changed as the molecular systematics project progressed. For an up to date publication on nomenclature and the new names and combinations, see our recent paper cited above.

Subtribe Maxillariinae: Proposed Species Alliances

1J. T. Atwood, 2G. Carnevali, 3C. H. Dodson

The long list below was an attempt by several workers to bring together and organize all the names associated with the genus Maxillaria into a purposely flexible classification borrowing heavily from the work of Pabst and Dungs (Orchidaceae Brasilienses, volume 2). The list attempts to do no more than to bring together likes with likes, and no categories are formal. Questions of relationships are provided and names for which we have almost no information are indicated as "Alliance Unknown" at the end of Maxillaria. No attempt is made to compare the system of Senghas (in R. Schechter, Die Orchideen, third edition) , although comparison will no doubt stimulate lengthy discussion. This list is a product of the digestion of over five hundred original descriptions and any available line drawings, photographs or specimens, types and otherwise seen by the describing authors of the various species. Names in parenthesis following a formal entry justified to the left include hometypic synonyms and possible heterotypic synonyms, but most of the latter are at present mostly matters of opinion. A more complete list of species can be found by searching for Maxillaria on the Kew web site World Checklist of Monocotyledons. Eric Christenson (2002) has published an overview of Maxillaria in the Proceedings of the 16th World Orchid Conference pp. 279-290, where he proposes a number of infrageneric divisions (see Literature for a complete citation). His work appeared in print well after this web site was first constructed, but we thank him nevertheless for pointing out that we did not forsee his publication. He maintains Sepalosaccus as a distinct genus, but includes Ornithidium in Maxillaria. Our molecular work is completely at odds with this proposal. Any further discussion of subgeneric divisions of the genus awaits additional molecular and morphological work. We regret any omissions and errors and we will attempt to correct them as work progresses. We always appreciate polite, constructive criticisms. We do not listen to whinning or shouting.

Most of Atwood's specimens are in TROPICOS.


Anthosiphon roseans Schltr. (=Cryptocentrum roseans (Schltr.) A.D.Hawkes)
Anthosiphon sp. (Costa Rica, or Panama)
Our molecular work supports lumping this genus into Cryptocentrum.

CHRYSOCYCNIS Lind. & Rchb.f.

C. ecuadorense Dodson & Garay
C. lehmannii Rolfe
C. schlimii Linden & Rchb.f. (Chrysocycnis trigonidii Lind. & Rchb.f.)
Hypothesis: Maxillaria tigrina C. Schweinf. belongs here despite atypical pollinia.

Excluded Taxa
Chrysocycnis glumaceum L.O.Williams (Cyrtidiorchis alatum (Lindl.) Rauschert
C. rhomboglossum Lehm. & Kraenzl. (Cyrtidiorchis rhomboglossum (Lehm. & Kraenzl.) Rauschert
C. tripterum Schltr. (Cyrtidiorchis rhomboglossum (Lehm. & Kraenzl. Rauschert,
(Cyrtidiorchis triptera (Schltr.) Rauschert)


The classification given here is based upon the doctoral dissertation of Germán Carnevali, and is not likely to change.

Cryptocentrum subgenus Cryptocentrum

C. caespitosum Carnevali (ined.)
C. calcaratum (Schltr.) Schltr. (=Pittierierella calcarata Schltr.)
C. dodsonii Carnevali (ined.)
C. dunstervilleorum Carnevali & G.A.Romero (C. gracillimum of authors, not Ames & C.Schweinf.)
C. escobarii Carnevali (ined.)
C. gracillimum Ames & C.Schweinf.
C. hirtzii Dodson
C. inaequisepalum C.Schweinf. (C. jamesonii of authors, not Benth.)
C. latifolium Schltr. (C. latifolium var. brachypetalum Garay; C. chocoensis Hawkes, name only)
C. lehmannii (Rchb.f.) Garay (Aeranthus lehmannii Rchb.f.; C. gracilipes Schltr.;C. jamesonii Benth.)
C. longipetalum Carnevali (ined.)
C. silverstonei Carnevali (ined.)
C. spathaceum Dodson

Cryptocentrum subgenus Polysticha Carnevali

C. flavum Schltr. (C. pervuianum of authors, not (Cogn.) C.Schweinf.
C. peruvianum (Cogn.) C.Schweinf. ssp. peruvianum (=Centroglossa peruviana Cogn.; C. hoppii Schltr.)
C. peruvianum ssp. dactylinum Carnevali, ined.
C. peruvianum ssp. minus (Schltr.) Carnevali, ined.(=C. minus Schltr.)
C. standleyii Ames (C. longiscapum Brieger; C. brenesii, name only)

Cryptocentrum subgenus Pseudobulbosa Carnevali

C. pseudobulbosum C.Schweinf.

Excluded names

C. costaricensis (Schltr.), comb. ined. (=Pittierella costaricensis Schltr., invalid)
C. pergracile Schltr. (no type material has been found)

CYRTIDIORCHIS Hypothesis: belongs with Chrysocycnis
(=Cyrtidium Schltr. 1924 not Vainio, 1921)

C. alata (Lindl.) Rausch. (=Camaridium alatum Lindl., Cyrtidium alatum (Lindl.) Garay; Chrysocycnis glumaceum L.O.Williams)
C. frontinoensis (Garay) Rauschert
C. rhomboglossa (Lehm. & Kraenzl.) Rauschert (Chrysocycnis rhomboglossa Lehm. & Kraenzl.,
Cyrtidium rhomboglossum (Lehm. & Kraenzl.) Schltr.; Cyrtidiorchis triptera (Schltr.) Rausch.,
Chrysocycnis tripterum Schltr., Cyrtidium tripterum (Schltr.) Schltr.)
C. stumpflei (Garay) Rauschert (=Cyrtidium stumpflei Garay)


Acuminata Alliance Hypothesis: close to Friedrichsthalii Alliance

M. acuminata Lindl.
M. cerifera Barb.Rodr. (=Ornithidium ceriferum (Barb.Rodr.) Barb. Rodr.; M. divaricata (Barb.Rodr.) Cogn., M. nervosa Rolfe ex Britton, M. flavoviridis Barb.Rodr. Ornithidium flavoviride (Barb.Rodr.) Barb.Rodr., Ornithidium divaricatum Barb.Rodr.)
M. huancabambae (Kraenzl.) C.Schweinf. (=Ornithidium huancabambae Kraenzl.; M. huancabambae (Kraenzl.) C.Schweinf. var. cuzcoensis C.Schweinf.
M. mariaisabelae J.T.Atwood
M. microblephara Schltr.
M. monacensis Kraenzl.
M. neillii Dodson [Pseudobulbs with three, linear, apical leaves.]
M. notylioglossa Rchb.f. (M. meirax Rchb.f. & Warm.; ?=M. fallax Schltr. fide C. Schweinfurth)
M. ophiodens J.T.Atwood
M. polybulbon Kraenzl.
M. ramosissima Kraenzl.
M. schistostele Schltr. [only 1 leaf at pseudobulb apex]
M. sulfurea Schltr. [Flowers sulfur yellow.]
M. unguiculata Schltr.
M. unguilabia Schltr.

Alba Alliance (abscission layers, but no blades at pb base, colors white to green to purplish or brownish?)

M. alba (J.D.Hook.) Lindl. (=Dendrobium album Hook.f., Broughtonia alba (Hook.f.) Spreng.,
Camaridium album (Hook.f.) Hoehne, ?=M. hedyosma Schltr., ?M. lactea Schltr. 1925, not 1923)
M. anceps Ames & C.Schweinf.
M. diuturna Ames & C.Schweinf.
M. nagelii L.O.Williams ex Correll (=Camaridium tuerckheimii Schltr.)
M. oreocharis Schltr.
[?Camaridium caquetanum Schltr.]

Albata Alliance

M. albata Lindl. [Carnevali has this as isolated]

Alpestris Alliance [Is this near Trigonidium as suggested by Carnevali? Or near Mormolyca?]

M. alpestris Lindl. (?=Maxillaria baumanniana Schltr., M. chlorochila Lehm. & Kraenzl. ex Kraenzl.,
M. compressibulba Schltr., M. trigona, C.Schweinf., M. xantholeuca Schltr., M. xantholeuca var. peruviana C.Schweinf., Maxillaria peruviana (C.Schweinf.) D.E.Benn. & Christenson)
M. bomboizensis Dodson
M. brachypetala Schltr.
M. casta Kraenzl. [flowers white]
M. chlorochila Lehm. & Kraenzl. ex Kraenzl.
M. cornuta C.Schweinf.
[Mormolyca fuchsii J.T.Atwood?]
M. koehleri Schltr. (?=M. laevilabris Lindl.)
M. laevilabris Lindl.
[M. mathewsii of authors, not Lindl. (1845) nor Rchb.f. (1863)]
M. napoensis Dodson (long foot)
M. piestopus Schltr.
M. sessilis Lindl., not M. sessilis (Sw.) Fawc. & Rendle
M. unicarinata C.Schweinf.

Hastulata/Pulla Suballiance (poorly developed column foot)

[Mormolyca fuchsii J.T.Atwood belongs here?]
M. dalstromii Dodson (=pulla?)
M. hastulata Lindl.
M. longipes Lindl. [possibly belongs with Parkeri group]
M. macrura Rchb.f. (Syn.: M. longisepala Rolfe)
M. nutantiflora Schltr.? (No types found, but description of plant and illustration of flower seems to fit here)
M. pannieri Fold.
M. pulla Lind. & Rchb.f.
M. rotundilabia C.Schweinf.

Grobyoides Suballiance (No foliaceous sheaths; pseudobulbs 2-foliate. Near M. picta? (Trigonidium?)

M. grobyoides Garay & Dunsterville
M. steyermarkii Fold.

Biolleyi Alliance

Amabilis Suballiance

M. amabilis J.T.Atwood
M. monteverdensis J.T.Atwood & G. Barboza
M. synsepala J.T.Atwood
M. tricarinata J.T.Atwood

Biolleyi Suballiance

M. adolphii Ames & Correll (=Camaridium adolphii Schltr., Ornithidium tonduzii Schltr.)
M. biolleyi (Schltr.) L.O.Williams (=Ornithidium biolleyi Schltr. Camaridium biolleyi (Schltr.) Schltr.)
M. falcata Ames & Correll (=Ornithidium costaricense Schltr.)
M. quadrata Ames & Correll (=Ornithidium lankesteri Ames)
M. tutae J.T.Atwood

Inaudita Suballiance

M. inaudita Rchb.f.
M. inaudita var. minor Ames & C.Schweinf.
?[trilobata Ames & C.Schweinf. Suballiance]
M. trilobata A. & C., (=Camaridium brenesii Schltr.)

Scalariformis Suballiance [This may bridge the gap between Biolleyi and Camaridium alliances. Lip is similar to that of M. bradeorum but monopodial canes belongs in Biolleyi Alliance.]

M. scalariformis J.T.Atwood

Tonduzii Suballiance

M. parvilabia Ames & C.Schweinf.
M. tonduzii (Schltr.) Ames & Correll (=Camaridium tonduzii Schltr., Camaridium costaricense Schltr.,
Maxillaria heidelbergensis Senghas, nom. illeg.)

Camaridii Alliance

Bradeorum Suballiance

M. ampliflora C.Schweinf. (Camaridium grandiflorum Ames, not Schltr.)
M. bradeorum (Schltr.) L.O.Williams (=Camaridium bradeorum Schltr; M. semiorbicularis Ames & C.Schweinf.)
M. campanulata C.Schweinf.
M. carinulata Rchb.f. [close to M. ampliflora]
M. darienensis J.T.Atwood
M. lamprochlamys (Schltr.) [comb. ined.] (=Camaridium lamprochlamys Schltr.)
M. umbratilis L.O.Williams (=Camaridium nutantiflorum Schltr., C. vinosum Schltr., M. vinosa (Schltr.) Senghas)

Camaridii Suballiance

M. camaridii Rchb.f. (=Cameridium ochroleucum Lindl., Cymbidium ochroleucum Lindl.;
?=Ornithidium album Hook., Maxillaria lutescens Scheidw., Camaridium lutescens (Scheidw.)
Rchb.f., Ornithidium fragrans Rolfe, Camaridium cryptodanthum of authors, Camaridium.
cyrtopodanthum Barb. Rodr., Camaridium affine Schltr., Camaridium amazonicium Schltr., Maxillaria hoehneana P.F.Hunt)
M. cozieriana H.G.Jones?????
M. suaveolens Barringer
M. pulchra (Schltr.) L.O.Williams ex Correll
M. insolita Dressler?????

Chicana Alliance [Only a single species here that is unlike any others. Plants have ovoid pseudobulbs with a single terete apical leaf. Lip has well defined lateral lobes above the middle.]

M. chicana Dodson

Ctenostachya Alliance

M. ctenostachya Rchb.f. (=Camaridium arachnites Schltr.)

Cucullata Alliance (close to Imbricata Alliance?)

M. atrata Rchb.f. (=M. soconuscana D.Breedlove?)
M. cacaoensis J.T.Atwood
M. cedralensis J.T.Atwood & D.E.Mora
M. cucullata Lindley
M. hematoglossa Rich. & Gal.)
[M. herzogiana Kraenzl., possibly belongs in Alpestris or Rufescens Alliance.]
M. lexarzana Soto Arenas & F.Chiang (=Psittacoglossum atratum Llave & Lex.)
M. lindeniana Rich. & Gal.
M. meleagris Lindl.
M. mombachoensis Heller ex J.T.Atwood
M. obscura Lind. & Rchb.f. (?=M. atrata var. brachyantha Rchb.f.)
M. praestans Rchb.f.
M. punctostriata Rchb.f. (=M. rubrilabia Schltr.)
M. rhombea Lindl.
M. seymouriana J.T.Atwood

Densa Alliance [Near Cucullata Alliance?]

M. densa Lindl. (=Ornithidium densum (Lindl.) Rchb.f., ?Maxillaria glomerata Gal.)
M. hagsateriana Soto Arenas

Densifolia Alliance (with or without pseudobulbs, always forming elongate canes, sheaths foliaceous, flowers with short, broad segments)

Densifolia Suballiance ["Broad" leaves]

M. alfaroi Ames & C.Schweinf.
M. cuencana Garay (=Ornithidium luteorubrum Lehm. & Kraenzl., Camaridium luteorubrum (Lehm. & Kraenzl.) Schltr.)
M. densifolia (Poepp. & Endl.) Rchb.f. (=Dicrypta densifolia Poepp. & Endl. 1835, Camaridium densifolium (Poepp. & Endl.) Benth. & Hook. 1883) [are pseudobulbs really absent as in original description?]
M. funicaulis C.Schweinf.
M. guareimensis Rchb.f.
M. johannis Pabst (=Camaridium robustum Barb.Rodr.)
M. platypetala Ruiz & Pav. (=Dendrobium platypetalum (Ruiz & Pav.) Pers., note specimen at K;?
M. polypetala Steud. (Schweinfurth suggests M. platypetala may be a Lycaste.)
M. procurrens Lindl.
M. saxatilis Rchb.f. 1855
M. schnitteri Schltr.
M. spilotantha Rchb.f.
M. subpandurata Schltr. [Schlechter compares this to M. variabilis which lacks foliaceous sheaths.]
M. xanthorhoda Schltr.[Description needed]

Arbuscula Suballiance ["Narrow" leaves]

M. arbuscula Rchb.f., not Camaridium arbuscula Lindl. (?=Fernandezia punctata Ruiz & Pav.)
M. arbuscula (Lindl.) comb. ined. (=Camaridium arbuscula Lindl.)
M. camaridioides Schltr.
M. christobalensis Rchb.f. (=Camaridium christobalense (Rchb.f.) Hoehne)
M. lawrenceana (Rolfe) Garay & Dunsterv. (=Camaridium lawrenceanum Rolfe)
M. microdendron Schltr.
?M. pamplonensis Lind. & Rchb.f.
M. pastensis Rchb.f. [type needed]
M. purpurata (Lindl.) Rchb.f. (=Camaridium purpuratum Lindl.)
M. stenophylla Rchb.f. (1854) not Lehm. & Kraenzl. (1899)
M. stictantha Schltr.
?M. vulcanica Lehm. & Kraenzl.

Ponerantha Suballiance

M. acervata Rchb.f. (=M. foliosa Ames & C.Schweinf.)
M. ponerantha Rchb.f.
M. swartziana C.D.Adams (=Epidendrum proliferum Sw., Cymbidium proliferum (Sw.) Sw., Isochilus prolifer (Sw.) R.Br., Ornithidium proliferum (Sw.) Fawc. & Rendle)
M. caucae Garay (=M. parvula Schltr.) (?=M. ponerantha of authors, not Rchb.f. (Purple flowered. Possibly allogamous forms of M. swartziana)
[Camaridium] vinosum Rolfe, not Schltr. Flowers are described as wine colored. This may be an allogamous form of M. swartziana.

Desvauxiana Alliance

M. auyantepuiensis Foldats
M. brachypoda Schltr.
[M. caparaoensis Brade]
M. desvauxiana Rchb.f. (=M. coriacea Barb. Rodr., M. petiolaris A. Rich. ex Rchb.f., M. verrucifera C.Schweinf.)
M. foldatsiana Carnevali & Ramirez
M. huebneri Schltr.
M. longipetiolata Ames & C.Schweinf.
M. riopalenquensis Dodson (=M. auyantepuiensis ssp. epiphytica Carnevali & Ramirez) [not M. neophylla Rchb.f.]

Dicrypta (Heterotaxis) Alliance (including Valenzuelana Suballiance of Pabst)
[M. boliviensis Schltr.?]

Bicallosa Suballiance (Leaves thin, never monopodial)

M. bicallosa (Rchb.f.) Garay (=Zygopetalum bicallosum Rchb.f., Kefersteinia bicallosa (Rchb.f.) Rchb.f., Ornithidium dolichophyllum Schltr., M. caespitosa C.Schweinf., Trigonidium equitans Garay)
M. bolleoides Schltr. (Possibly the same as M. chartacifolia)
M. chartacifolia Ames & C.Schweinf.
M. sp. nov., "spotted"
Ornithidium cyperifolium Schltr. [close to M. bicallosa; flowers "wine red"]
Ornithidium rhodoleucum Schltr. [close to M. bicallosa; flowers dark red with white lip]

Crassifolia Suballiance

[M. brasiliensis Brieger & Bicalho?]
M. crassifolia (Lindl.) Rchb.f. (=Heterotaxis crassifolia Lindl., Dicrypta crassifolia (Lindl.) Lindl. ex Loud., Epidendrum sessile Sw., M. sessilis (Sw.) Fawc. & Rendle, Dicrypta baueri Lindl.; ?=M. gatunensis Schltr.)
M. cymbidioides Dodson, J.T.Atwood, & Carnevali
M. discolor (Lodd. ex Lindl.) Rchb.f. (=Dicrypta discolor Lodd. ex Lindl.)
M. maleolens Schltr.
M. nasuta Rchb.f. (=M. nasalis Rchb.f., M. brevipedunculata Ames & C.Schweinf., ? M. oxysepala Schltr.)
M. proboscidea Rchb.f.
M. santanae Carnevali & Ramirez
M. superflua Rchb.f. (M. longifolia (Barb.Rodr.) Cogn. not Lindl., M. tarumaensis Hoehne)
M. verecunda Schltr. [close to M. villosa?]
M. villosa (Barb.Rodr.) Cogn. (=Dicrypta villosa Barb. Rodr.)
M. violaceopunctata Rchb.f.

Valenzuelana Suballiance

M. equitans (Schltr.) Garay (=Camaridium equitans Schltr., M. mattogrossensis Brade, Camaridium vandiforme Schltr., Marsupiaria vandiformis (Schltr.) Hoehne, M. vandiformis (Schltr.) C.Schweinf.)
M. fritzii Carnevali & Dodson, ined.
M. valenzuelana (A.Rich) Nash (=Pleurothallis valenzuelana A.Rich., Marsupiaria valenzuelana (A.Rich) Garay, Dicrypta iridifolia Bateman ex Rchb.f., M. iridifolia (Batem. ex Rchb.f.) Rchb.f.,
Marsupiaria iridifolia Hoehne, Dicrypta irisphyta Barb.Rodr., M. valenzuelana ssp. angustata J.T.Atwood
M. witsenioides Schltr.

Elatior Alliance (close to Densifolia Alliance?)

M. elatior (Rchb.f.) Rchb.f. (=Dicrypta elatior Rchb.f., M. triangularis Lindl., M. albertii Schltr.; ?=Camaridium xylobiichilum Kraenzl.)
M. tubercularis J.T.Atwood

Floribunda Alliance

M. caveroi D.E.Benn. & Christenson
M. dodsonii Carnevali, ined.
M. floribunda Lindl.
M. merana Dodson
M. platyloba Schltr. (Possibly misapplied to this alliance)
M. quelchii Rolfe
M. dunstervillei Carnevali & Ramirez
M. yanganensis Dodson

Friedrichsthalii Alliance (Hypothesis: close to Acuminata Alliance)

M. aciantha Rchb.f. (=Lycaste aciantha (Rchb.f.) Rchb.f.)
M. encyclioides Dodson & J.T.Atwood
M. friedrichsthalii Rchb.f. (=M. turialbae Schltr.)
[?M. nymphopolitana Kraenzl.]
M. ochroglossa Schltr.
M. scorpioidea Kraenzl. (=Camaridium scorpioideum (Kraenzl.) Hoehne; ?=Maxillaria rhodosticta Kraenzl.)

Graminifolia Alliance (may be able to divide into three or more suballiances)

Dendrobioides suballiance

M. allenii L.O.Williams
M. dendrobioides (Schltr.) L.O.Williams (=Camaridium dendrobioides Schltr., Camaridium jimenezii Schltr., ?Camaridium simile Schltr.)
M. valerioi Ames & C.Schweinf.
M. fragrans J.T.Atwood

Graminifolia Suballiance : No pseudobulbs in adults, lanceolate leaves

M. cassapensis Rchb.f. (=Camaridium cassapense (Rchb.f.) Schltr.
M. graminifolia (Kunth) Rchb.f. (=Isochilus graminifolius Kunth, Camaridium graminifolium (Kunth) Rchb.f.). M. lancifolia (Rchb.f.) Rchb.f. (=Camaridium lancifolium Rchb.f.) M. polyphylla Rchb.f. (=Camaridium polyphyllum (Rchb.f.) Schltr.). M. similis Garay & Dunsterville (=M. mathewsii Rchb.f. (1863) not Lindl. (1845), Camaridium mathewsii (Rchb.f.) Schltr.)

M. linearifolia Ames & C.Schweinf. (very close to M. casapensis, but has longer ovary and is from Costa Rica and Panama.)

Luteorubra Suballiance: No pseudobulbs in adults, elliptic leaves

M. appendiculoides C.Schweinf.
M. brevifolia (Lindl.) Rchb.f. (=Camaridium brevifolium Lindl.)
M. caucana (Schltr.) [needs new name?] (=Camaridium caucanum Schltr.)
M. flavum (Schltr.), unpublished homonym (=Camaridium flavum Schltr.)
M. haenkei Correll (=Isochilus grandiflorum Lindl., M. longibracteata (Lindl.) Rchb.f. var. grandiflora (Lindl.) C.Schweinf.
M. longibracteata (Lindl.) Rchb.f. (=Camaridium longibracteatum Lindl.)
M. luteorubra (Lindl.) Rchb.f. (=Camaridium luteorubrum Lindl.)
M. nitidula Rchb.f.

Suballiance in question: pseudobulbs present in mature shoots

M. arbuscula Rchb.f.
M. sanguineomaculata Schltr.
M. sodiroi (Schltr.) Dodson (=Camaridium sodiroi Schltr.)

Grandiflora Alliance (Close to Lepidota Alliance?)

This group interfaces with both Lepidota and Parkeri groups and may not be separable from them. The Grandiflora Alliance is characterized by the plant habit-cespitose with apically 1-foliate leaves, subtended by 1-2 foliaceous sheaths, and the the flowers which are generally large for the genus, at least with broad perianth parts and lip with pseudopollen. Flower colors are usually white, less often tinged yellow, sometimes spotted with large pokadots. Plants are mostly cool wet growing mid to high Andean plants that are impossible to grow in hot climates.

M. amesiana Hort. ex Gard. Chron.
M. anatomorum Rchb.f.
M. augustaevictoriae Lehm. & Kraenzl. ex Kraenzl. (syn.: M. hirtzii Dodson & A.Andreetta)
M. dalessandroi Dodson
M. dichroma Rolfe
M. eburnea Lindl.
M. elegantula Rolfe [Yellowish and spotted apically]
M. fletcheriana Rolfe
M. fucata Rchb.f.
M. grandiflora (HBK) Lindl.
M. grandis Rchb.f.
M. grayi Dodson [Or could this possibly be close to the Bicallosa Suballiance?]
M. hillsii Dodson
M. hoppii Schltr.
M. huebschii Rchb.f. [Possibly same as M. fucata Rchb.f.]
M. insignis Rolfe
M. irrorata Rchb.f.
M. johniana Kraenzl.
M. jucunda Lehm. & Kraenzl.
M. kalbreyeri Rchb.f.
M. lehmannii Rchb.f.
M. lindenii Cogn.
M. litensis Dodson
M. melina Lindl.
M. mirabilis Cogn. [yellow base color]
M. molitor Rchb.f. [yellow base color]
M. mooreana Rolfe
M. nutans Lindl.
M. sanderiana Rchb.f.
M. striata Rolfe
M. urbaniana Lehm. & Kraenzl.
M. venusta Lind. & Rchb.f.


M. nuriensis Carnevali & Ramirez

Imbricata Alliance

M. bracteata (Schltr.) Ames & Correll (=Ornithidium bracteatum Schltr., Camaridium bracteatum (Schltr.)Schltr.; M. vagans Ames & C.Schweinf.)
M. gomeziana J.T.Atwood (?=M. admonens Bock)
M. imbricata Barb.Rodr. (Camaridium imbricatum (Barb.Rodr.) Hoehne 1947 non Schltr. 1918, M. iguapensis Hoehne & Schltr., Camaridium iguapense (Hoehne & Schltr.) Hoehne, Camaridium imbricatum (Barb.Rodr.) Hoehne, Maxillaria carinata Barb. Rodr., Camaridium carinatum (Barb.Rodr.) Hoehne, Camaridium hoehnei Pabst)
M. praetexta Rchb.f. [1854, probably an earlier name for M. imbricata Barb. Rodr. (1877)]

Lepidota Alliance

Lepidota Suballiance

[M. angustissima Ames, Hubbard, & C.Schweinf. (=M. acutifolia Schltr., not Lindl.)]
M. arachnites Rchb.f.
M. arachnitiflora Ames & C.Schweinf.
M. attenuata Ames & C.Schweinf. (?=M. guadalupensis)
M. bolivarensis C.Schweinf.
M. bradei Schltr. ex Hoehne
M. caucana Schltr., not Camaridium caucanum Schltr.
M. colorata Rchb.f.
M. confusa Ames & C.Schweinf.
M. crocea Lindl. not Poepp. & Endl. (=M. lindleyana Schltr.)
M. cuzcoensis C.Schweinf.
M. ecuadorensis Schltr.
M. edwardsii D.E.Benn. & Christenson
[?M. elegans Schltr.; close to M. reichenheimia fide Schltr.]
M. farinifera Schltr.
M. fractiflexa Rchb.f.
M. guadalupensis Cogn.
M. guiardiana Chiron
M. lepidota Lindl. (M. pertusa Lindl. ex Rchb.f., M. saxicola Schltr.)
M. lilliputana D.E.Benn. & Christenson
M. longissima Lindl. (M. pentura Lindl.)
M. macropoda Schltr.
M. marmoliana Dodson
M. muscicola Rchb.f., 1878 (=earlier name for M. arachnites Rchb.f., 1880?)
[?M. neophylla Rchb.f.]
M. chionantha J.T.Atwood
M. pachyacron Schltr.
M. patens Schltr.
M. pseudoreichenheimiana Dodson
M. ramonensis Schltr. (=M. brevipes Schltr.)
M. reichenheimiana Endr. & Rchb.f.
M. rubioi Dodson
M. simplicilabia C.Schweinf.
M. thurstoniorum Dodson
M. trilobulata D.E.Benn. & Christenson
M. virguncula Rchb.f.
M. whittenii Dodson

Pterocarpa Suballiance

M. brachybulbon Schltr. (1-2 lvs. at base of bulb)
[M. liparophylla Summerhayes (This species is compared by the author to a large M. pterocarpa but may not even belong to the Suballiance)]
M. pterocarpa Barb.Rodr.
[M. xylobiiflora Schltr.]

Speciosa Suballiance (very close to Lepidota Suballiance)

M. gentryi Dodson
M. langlassei Schltr.
M. rodrigueziana J.T.Atwood & Mora
M. speciosa Rchb.f. (=M. scurrilis Rolfe)

Splendens Suballiance (three or more leaves at base of pseudobulb, flowers with linear tepals)

M. chlorantha Lindl.
M. gracilipes Schltr.
M. granditenuis D.E.Benn. & Christenson
M. klugii C.Schweinf.
M. mathewsii Lindl., not Rchb.f.
M. ochroleuca Lodd. ex Lindl.
M. ochroleuca var. longipes Sander
M. pauciflora Barb.Rodr.
M. rodriguesii Cogn. (=M. longipetala Barb. Rodr., not Ruiz Pav.)
M. splendens Poepp. & Endl. (M. histiologorum Rchb.f., fide Pabst, ? M. buchtienii Schltr.)
M. tenuis C.Schweinf.
M. weberbaueri Schltr.

Suballiance unknown

M. bennettii Christenson [Possibly this belongs in its own alliance]
M. hirsutilabia D.E.Benn. & Christenson

Lueri Alliance (Germán says this may be part of Meridensis Alliance)

M. lueri Dodson
M. linearis C.Schweinf.

Meridensis Alliance

[?M. caquetana Schltr.]
[?M. caulina Schltr.]
[M. chasei Dodson]
[?M. elata Schltr.]
M. embreei Dodson
M. exaltata (Kraenzl.) C. Schweinf. (=Camaridium exaltatum Kraenzl.)
M. longicaulis Schltr.
M. meridensis Lindl.
M. nanegalensis Rchb.f.
M. schlechteri Foldats (=M. rugosa Schltr. 1914, not Scheidw. 1843; Camaridium rugosum (Schltr.) Hoehne; M. sulcata C.Schweinf., illeg.
M. williamsii Dodson

Microphyton Alliance

M. burgeri J.T.Atwood
M. flava Ames, Hubbard, & C.Schweinf. (=Ornithidium ramonense Schltr.)

Wercklei Suballiance

M. jugata Garay
M. lankesteri Ames (=Ornithidium aurantiacum Schltr.)
M. microphyton Schltr. (=O. parvulum Schltr., O. pallidiflavum Schltr., M. pallidiflava (Schltr.) Senghas)
M. muscoides J.T.Atwood
M. oestlundiana L.O.Williams
M. wercklei (Schltr.) L.O.Williams (=Ornithidium wercklei Schltr.)

Nardoides Alliance Pseudobulbs apically 3-5 foliate. Interfaces with Pityphyllum?

M. nardoides Kraenzl.

Neourbania Alliance (close to or part of Ornithidium Alliance, Aurea Suballiance?)

M. adendrobium (Rchb.f.) Dressler (=Ponera adendrobium Rchb.f., Pleuranthium adendrobium (Rchb.f.) Benth. & Hook.f., Neourbania adendrobium (Rchb.f.) Fawc. & Rendle; Camaridium parviflorum Fawc.)
M. chrysocycnoides (Schltr.) Dodson. (=Ornithidium chrysocycnoides Schltr.)
M. conduplicata (Ames & C.Schweinf.) L.O.Williams (=Ornithidium conduplicatum Ames & C.Schweinf.)
M. haemathodes (Ruiz & Pav.) Garay (=Fernandezia haemathodes Ruiz & Pav.; Ornithidium weberbauerianum Kraenzl.; Maxillaria crassicaulis C.Schweinf.) [This is the now famous blue Maxillaria recently rediscovered in Bolivia.]
M. nicaraguensis (Hamer & Garay) J.T.Atwood (=Neourbania nicaraguensis Hamer & Garay)
M. machinazensis D.E.Benn. & Christenson [=M. mapiriensis?]
M. mapiriensis (Kraenzl.) L.O.Williams (=Ornithidium mapiriense Kraenzl., ?including Maxillaria condensata C.Schweinf.). Is this close to the Ramosa Alliance?
M. pastoensis (Schltr.) comb. ined. (=Ornithidium pastoense Schltr.)
M. pleuranthioides (Schltr.) Garay (=Ornithidium pleuranthioides Schltr.)
M. quitensis (Rchb.f.) C. Schweinf.
M. sp. nov. ined. identified as M. adendrobium (Rchb.f.) Dressler (Dodson 7364 (SEL); Dodson et al. 8511 (SEL), Ecuador

Ornithidium (Aggregata) Alliance (Neourbania Alliance may belong here)

M. affinis (Poepp. & Endl.) Garay (=Scaphyglottis affinis Poepp. & Endl., Ponera affinis Rchb.f.), not Camaridium affine Schltr., not M. affinis Hort. Petrop. ex Rchb.f.)
M. aggregata (H.B.K.) Lindl.
M. bicolor (Rolfe) (=Ornithidium bicolor Rolfe) Possibly this belongs in Biolleyi Alliance]
M. disticha (Lindl.) C.Schweinf. (Ornithidium distichum Lindl.)
M. jamesonii (Rchb.f.) Garay & C.Schweinf. (=Ornithidium jamesonii Rchb.f.)
?M. lehmannii (Kraenzl.) comb. ined.; needs new name, (Ornithidium lehmannii Kraenzl.), not Maxillaria lehmannii Rchb.f.
M. nubigena (Rchb.f.) C.Schweinf. (=Ornithidium nubigenum Rchb.f., ?=Pachyphyllum pamplonense Kraenzl.)
M. palmensis Dodson (=Ornithidium breve Schltr.) [Not Maxillaria brevis (Hoehne & Schltr.) Hoehne]
?M. pfitzerianum (Lehm. & Kraenzl.) comb. ined. (=Ornithidium pfitzerianum Lehm. & Kraenzl.)
M. squarrosa (Schltr.)?
M. torifera (Schltr.) comb. ined. (=Ornithidium toriferum Schltr.)
[Ornithidium] vagans Schltr.(not Maxillaria vagans Ames & C.Schweinf.)

Aurea Suballiance

M. aurea (Poepp. & Endl.) L.O.Williams (=Ornithidium aureum Poepp. & Endl., Camaridium aureum (Poepp. & Endl.) Hoehne)
Maxillaria cordyline (Rchb.f.) Dodson (=Ornithidium cordyline Rchb.f.)
M. crassicaulis C.Schweinf.
M. gigantea (Lindl.) Dodson (=Ornithidium giganteum Lindl., Camaridium aureum var. giganteum Hoehne; ?=Ornithidium bolivianum Schltr.)
M. gigantea ssp. condorensis D.E.Benn. & Christenson [this may be a good species, and close to M. conduplicata of Neourbania Alliance.]
M. latifolia (Kunth) Lindl. (fide C. H. Dodson) (=Dendrobium latifolium Kunth)
M. longa (Schltr.) Dodson (=Camaridium longum Schltr.)

Coccinea Suballiance

M. alticola C.Schweinf. (=Ornithidium serrulatum Lindl., not M. serrulata Ames & Correll)
M. coccinea (Jacq.) L.O.Williams ex Hodge (=Epidendrum coccineum Jacq., Ornithidium coccineum (Jacq.) Salisb., Camaridium coccineum (Jacq.) Hoehne
M. fulgens (Rchb.f.) L.O.Williams (=Ornithidium fulgens Rchb.f.)
M. heraldica Rchb.f.
M. jenischiana Rchb.f. (Ornithidium jenischianum Rchb.f.)
M. miniata (Lindl.) L.O.Williams (=Ornithidium miniatum Lindl.). Fls. crimson-vermillion; lip yellow edged and blotched with crimson.
M. nivea (Lindl.) L.O.Williams (=Ornithidium niveum Lindl.). Fls. white.
M. pittieri (Ames) L.O.Williams (Ornithidium pittieri Ames)
M. ruberrima (Lindl.) Garay (=Scaphyglottis ruberrima Lindl.). Fls. bright red.
M. sanguinolenta (Lindl.) C.Schweinf. (=Ornithidium sanguinolentum Lindl.). Fls. bright yellow; lip with red spot.
[Ornithidium] semiscabrum Lindl.

Paleata (wrightii) Alliance

M. dichotoma (Schltr.) L.O.Williams (=Camaridium dichotomum Schltr.)
?M. insolita Dressler
M. lutheri J.T.Atwood
M. minor (Schltr.) L.O.Williams (=Camaridium minus Schltr.)
M. paleata (Rchb.f.) Ames & Correll (=Ornithidium paleatum Rchb.f.; Camaridium wrightii Schltr., M. wrightii (Schltr.) Ames & Correll, Ornithidium wrightii (Schltr.) C.Schweinf.)
M. pardalina Garay (=M. pantherina Rchb.f. not Hoffmsgg.
M. schlechteriana J.T.Atwood (=Camaridium imbricatum Schltr., Ornithidium schlechterianum C.Schweinf. [nom. illeg.], M. wrightii (Schltr.) Ames & Correll var. imbricata (Schltr.) Ames & Correll.
M. serrulata Ames & Correll (=Camaridium amparoana Schltr., Ornithidium amparoanum (Schltr.) Pabst)
M. sigmoidea (C.Schweinf.) Ames & Correll (=Ornithidium sigmoideum C.Schweinf.)
M. vaginalis Rchlb.f. (=Camaridium wercklei Schltr.)

Parkeri Alliance [No foliaceous sheaths; pseudobulbs unifoliate]

M. aequiloba Schltr.
M. albiflora Ames & C.Schweinf.
M. angustisegmenta Ames, Hubbard, & C.Scweinf., Maxillaria endresii Rchb.f. var. angustisegmenta (Ames, Hubbard, & C.Schweinf.) C. Schweinf.)
M. argyrophylla Poepp. & Endl.
M. aurorae D.E.Benn. & Christenson
M. batemanii Poepp. & Endl. (may have a foliaceous sheath at pseudobulb base)
M. breviscapa Poepp. & Endl.
M. calantha Schltr.
M. callichroma Rchb.f.
M. caloglossa Rchb.f.
[M. candida Lodd. ex Lindl.]
M. dillonii D.E.Benn. & Christenson [close to M. callichroma Rchb.f.]
M. divitiflora Rchb.f.
M. endresii Rchb.f. (=M. leptosepala?)
M. galantha J.T.Atwood & Carnevali
M. grandimentum C.Schweinf.
M. lactea Schltr. 1925, not 1923 (close to M. yauaperiensis Barb.Rodr. and M. xanthosia Barb.Rodr.)
M. leptosepala Hook. (=M. robusta Barb.Rodr.?)
M. loretoensis C.Schweinf. (close to M. parkeri)
M. luteoalba Lindl.
M. monantha Barb.Rodr. (close to M. parkeri? Rufescens Alliance fide Pabst)
M. multiflora Barb.Rodr.(close to M. parkeri?)
M. nigrescens Lindl. (M. rubro-fusca Klotzsch)
?M. pachyneura Lehm. & Kraenzl.
M. parkeri (Spreng.) Hook. (=Colax parkeri Spreng., M. hirtilabia Lindl.?, M. lorifolia Rchb.f.?)
M. perryae Dodson
M. plicata Schltr.
?M. schultzei Schltr. (Flowers sulfur-yellow; lip painted with red. Schlechter believes this is close to M. leucaimata Barb.Rodr., but may be close to M. parkeri).
M. setigera Lindl.
M. shepheardii Rolfe (close to M. parkeri or M. schultzei?)
[M. taracuana Schltr., close to M. ringens?]
M. triloris Morren
M. verrucifera C.Schweinf.
M. woytkowskii C.Schweinf.
M. xanthosia Barb.Rodr.
M. yauaperyensis Barb.Rodr. [close to M. parkeri?]

Porrecta (Brunnea) Suballiance (lack of leaf at pb base, probably best as suballiance of Parkeri Alliance)

M. acostaei Schltr. (small M. brunnea-sort)
M. amazonica (?=M. powellii)
M. brunnea Lind. & Rchb.f. (including M. bogotensis Rchb.f. & Triana, name only based on W 51428, M. brenesii Schltr., and probably earlier described M. porrecta Lindl., M. trinitatis Ames, M. trinitensis Ames ex W.E.Broadway, name only)
[?M. aurantiaca Schltr., not M. aurantiaca A.Rich. & Gal. If a good species, a new name is needed]
M. cryptobulbon Carnevali & J.T.Atwood
M. dolichophylla Schltr.
M. furfuracea Scheidw. (belongs with M. porrecta fide Pabst)
[?M. glauca Lindl. ex W. Baxt. Description needed]
M. glumacea Rolfe (belongs with M. porrecta fide Pabst)
M. guentheriana Kraenzl. [Kraenzlin compares this with M. dolichophylla Schltr., but the pseudobulbs are described as apically bifoliate.]
M. kegelii Rchb.f. [or does this belong with M. ringens Suballiance?]
M. leucaimata Barb.Rodr.
M. leucantha Schltr. (1929), not Kl., Otto, & Dietr. (1850)
M. longiloba (Ames & C.Schweinf.) J.T.Atwood (=M. brenesii Schltr. var. longiloba Ames & C.Schweinf.
M. microtricha Schltr.
M. modestiflora Pabst (=M. modesta Brade. Close to M. cryptobulbon?)
M. pleiantha Schltr. [Schlechter compares this to M. melina Lindl.]
M. poicilothece Schltr. (close to M. brunnea)
M. porrecta Lindl. (M. aurea Hort. ex Rchb.f., M. mutabilis Hort. ex Rchb.f.) (includes M. brunnea?)
M. powellii Schltr. (part of M. brunnea or M. porrecta?)
[?M. simacoana Schltr.]
M. spiritu-sanctensis Pabst

Ringens Suballiance (close to Luteoalba Alliance?)

M. fuerstenbergiana Schltr.
M. hennisiana Schltr.
M. maderoi Schltr. [Schlechter compares this to M. lepidota but seems more similar to a large M. fuerstenbergiana.]
M. parvibulbosa C.Schweinf.
M. ringens Rchb.f. (?=M. amparoana Schltr., M. lactea Schltr. 1923 not 1925, M. pubilabia Schltr., M. rouseauae Schltr., M. tuerckheimii Schltr., M. yzabalana S.Watson, M. lactiflora Pabst, M. leucoxantha Endres, name only based on Endres collection at W)

Picta Alliance (Brazilian Alliance, two apical leaves)

M. brevis (Hoehne & Schltr.) Hoehne (=M. picta var. brevis Hoehne & Schltr.)
M. barbosae Loefgren ex Porto
M. consanguinea Kl. (M. picta Hook. var. brunnea Rchb.f., M. serotina Hoehne, not Regnell & Barb.Rodr.)
M. gracilis Lodd. (=M. acutipetala Hook., M. liliacea Barb. Rodr. not Ll. & Lex., M. penduliflora Fenzl.,
M. queirogana Barb.Rodr.)
[?M. guayanensis Klinge]
M. heismanniana Barb.Rodr.
M. kautskii Pabst
M. moutini Pabst (M. moutinhoi ex Index Kewensis)
?M. parviloba Rolfe [type needed. Origin was thought to have been Brazil.]
M. phoenicanthera Barb.Rodr. (M. punctata Schltr. not Lodd.)
M. picta Hook. (M. leucocheile Hoffmsgg., M. kreysigii [Regel?], M. fuscata Kl. ex Rchb.f. [1863], non Hort. ex Rchb.f. [1863], M. monoceros Kl. ex Rchb.f., M. pallidiflora Rchb.f. ex Barb. Rodr., illeg., not M. pallidiflora Hook.)
M. polyantha Barb.Rodr.
M. porphyrostele Rchb.f.
M. punctata Lodd.
M. regeliana Cogn. (M. galeottiana Regel, not Rich. et Gal.; Regel description of 1860 indicates "Mexico," but doubtful)
M. rupestris Barb.Rodr. (M. punctata Hoehne non Lodd., M. hoehnei Schltr.)
M. schunkeana Campacci & Kaustsky
[M. tristis Schltr.; but possibly close to M. grobyoides]
M. ubatubana Hoehne

Marginata Suballiance (elongate rhizomes)

M. chrysantha Barb.Rodr. (M. picta Rchb.f. non Hook., M. serotina Regnell & Barb.Rodr.)
M. marginata Fenzl. (M. tricolor Lindl., M. punctulata Kl., M. deflexa Kl. ex A.Br.Kl. & Bouche, M. crassipes Kraenzl., M. murilliana Hoehne)
M. piresiana Hoehne

Madida Suballiance (plants small, rarely pseudobulbs apically 1-foliate)

[?M. broadwayi (Cogn.) R.E.Schultes (=Ornithidium broadwayi Cogn.)]
M. cogniauxiana Hoehne, (=M. heterophylla Hoehne)
M. madida Lindl. (M. cepula Rchb.f., M. mosenii Kraenzl., M. hatschbachii (Schltr.) Hoehne, M. echinochila Kraenzlin)
M. neowiedii Rchb.f.
M. paranaensis Barb. Rodr.

Planicola Alliance

M. haberi J.T.Atwood
M. planicola C.Schweinf. (=Camaridium latifolium Schltr.)

Pseudomaxillaria Alliance

M. brevilabia Ames & Correll (Ornithidium alfaroi Ames & C.Schweinf.)
?M. compacta (Schltr.) comb. ined. (=Ornithidium compactum Schltr.) [The flower is most similar to that of M. parviflora, but description of plant habit is very different.]
M. concavilabia Ames & Correll (Ornithidium stenophyllum Schltr.)
M. horichii Senghas (=Sepalosaccus horichii (Senghas) Senghas)
M. neglecta (Schltr.) L.O.Williams (=Ornithidium neglectum Schltr.)
M. parviflora (Poepp. & Endl.) Garay (=Scaphyglottis parviflora Poeppig & Endl. Ornithidium parviflorum (Poepp. & Endl.) Rchb.f., Pseudomaxillaria parviflora (Poepp. & Endl.) Brieger;
Ornithidium confertum Griseb., M. conferta Ames & C.Schweinf., Ornithidium chloroleucum Barb. Rodr., Pseudomaxillaria chloroleuca (Barb.Rodr.) Hoehne, M. simulans (Ames & C.Schweinf.) L.O.Williams, Ornithidium simulans Ames & C.Schweinf., M. exigua Regel, M. surinamensis Focke ex Rchb.f., M. purpurea Ames & Correll, Ornithidium rhomboglossum Schltr., Pseudomaxillaria rhomboglossa (Schltr.) Brieger, Epidendrum vestitum Sw., Cymbidium vestitum (Sw.) Sw., Camaridium vestitum (Sw.) Lindl., Pseudomaxillaria vestita (Sw.) Sw. Brieger, Ornithidium vestitum(Sw.) Rchb.f., Camaridium purpureum Spreng.)
M. pseudoneglecta J.T.Atwood (=Ornithidium anceps Rchb.f., Ornithidium xanthinum Rchb.f., ined.)
M. virescens (Schltr.) comb. ined. (=Ornithidium virescens Schltr., Pseudomaxillaria virescens (Schltr.) Brieger

Pumila Alliance (including Paulistana Alliance. Always a single leaf, but close to or perhaps belonging to Madida suballiance of Picta Alliance)

M. ferdinandiana Barb.Rodr
M. fuscopurpurea Drapiez
M. funerea Lindl.
M. minuta Cogn.
M. pachyphylla Schltr. ex Hoehne
M. parahybunensis Cogn.
M. parva Rolfe
M. paulistana Hoehne
M. plebeja Rchb.f.
M. pumila Hook.
?M. pusilla Rolfe
M. spannagelii Hoehne

Ramosa Alliance [Related to Neorbania group?]

M. fasciculata C.Schweinf.
M. histrionica (Rchb.f.) L.O.Williams (=Ornithidium histrionicum Rchb.f., Maxillaria aristeguietae Foldats) [?M. inflexa (Lindl.) Griseb. (=Ornithidium inflexum (Lindl.) Rchb.f., Camaridium inflexum Lindl.; ?=Ornithidium croceorubens Rchb.f., Maxillaria croceorubens (Rchb.f.) L.O.Williams]
M. lasallei Fold.
M. loefgrenii (Cogn.) Pabst (=Ornithidium loefgrenii Cogn., Camaridium loefgrenii (Cogn.) Hoehne,
M. spathulata C.Schweinf.)
M. multicaulis (Poepp. & Endl.) C.Schweinf. (=Siagonanthus multicaulis Poepp. & Endl., Ornithidium multicaule (Poepp. & Endl.) Rchb.f., Camaridium multicaule (Poepp. & Endl.) Hoehne)
M. ochracea (Rchb.f.) Garay (=Ornithidium ochraceum Rchb.f.)
M. pendens Pabst (=Camaridium pendulum Barb.Rodr. 1877)
M. pendula (Poepp. & Endl.) C.Schweinf. (=Scaphyglottis pendula Poepp. & Endl., Ornithidium pendulum cogn.)
[M. perijaensis Carnevali & Romero, G. ined.]
M. ramosa Ruiz & Pav. (Dendrobium ramosum Pers., ?=Scaphyglottis taphallae Rchb.f., Ornithidium taphallae (Rchb.f.) Rchb.f., Maxillaria taphallae (Rchb.f.) C.Schweinf., O. dichotomum Schltr.)
M. repens L.O.Williams
M. rigida Barb. Rodr. (=Camaridium rigidum Schltr.)
M. sillarensis Dodson & Vásquez
M. tonsoniae Soto Arenas
[Camaridium sterrocaulon Schltr.]
[Camaridium vagans Schltr.], not Maxillaria vagans Ames & C.Schweinf.

Rufescens Alliance (hypothesis: Mormolyca ringens (Lindl.) Schltr. belongs here)

M. abelei Schltr.
M. acutifolia Lindl., not Schltr.
M. articulada Kl., fide Hammer, Icon. Pl. Trop. plate 1047
M. chacoensis Dodson
[M. cleistogama Brieger & Bicalho, fide Pabst]
M. dressleriana Carnevali & J.T.Atwood
M. hedwigiae Hamer & Dodson
M. moralesii Carnevali & J.T.Atwood
?M. muelleri Regel
M. richii Dodson
M. rufescens Lindl. (Syn.: M. fuscata Hort. ex Rchb.f. 1863, non Kl. ex Rchb.f.)
M. rugosa Scheidw.
M. suarezorum Dodson
M. vanillodora A.Rich & Gal.

Saragurensis Alliance (Near Nardoides Alliance or Pityphyllum?)

M. saragurensis Dodson. Odd plant with hooks on each side of the sheath apex.

Sepalosaccus Alliance (probably this belongs with the Parviflora Alliance)

M. strumata (Endres & Rchb.f.) Ames & Correll (=Ornithidium strumatum Endres & Rchb.f., Sepalosaccus strumatum (Endr. & Rchb.f.) Garay; ?=Sepalosaccus humilis Schltr.)

Sophronitis Alliance

M. jamboensis Dodson
M. patula C.Schweinf.
M. sophronitis (Rchb.f.) Garay (Ornithidium sophronitis Rchb.f.)

Subulata Alliance (close to Picta Alliance; leaves 2-3, terete or nearly so)

M. acicularis Herb.
M. echinophyta Barb.Rodr.
M. juergensii Schltr.
M. perpusilla Kraenzl.
M. poaefolia Schltr. (from Bolivia)
M. seidelii Pabst
M. subulata Lindl.
M. vernicosa Barb.Rodr.
M. vitelliniflora Barb.Rodr.

Tigrina Alliance (Hypothesis: close to or within genus Chrysocycnis)

M. tigrina C.Schweinf., (=Chrysocycnis tigrinum (C.Schweinf.) J.T.Atwood.)

Uncata Alliance

M. subulifolia Schltr.
M. uncata Lindl. (=Camaridium uncatum (Lindl.) Hoehne), ?M. macleei Batem. ex Lindl., ?M. nana Hook., ?M. squamata Barb.Rodr., Ornithidium squamatum (Barb. Rodr.) Barb. Rodr., Camaridium squamatum (Barb. Rodr.) Hoehne). ?M. stenostele Schltr., ?M. striatella Kraenzl.

Variabilis Alliance (Pseudobulbs apically 1-foliate; no foliaceous sheaths at base, flowers never with greatly attenuate floral segments.

M. adscendens Schltr.
M. angustifolia Hook. (?=M. variabilis Batem. ex Lindl.)
M. caespitifica Rchb.f. [?Syn.: M. modesta Schltr.]
M. cobanensis Schltr. [Carnevali has this near Pseudomaxillaria]
M. costaricensis Schltr.
M. estradae Dodson
M. houtteana Rchb.f.
M. infausta Rchb.f.
M. mexicana J.T.Atwood & Soto, ined.
M. panamensis Schltr.
M. sanguinea Rolfe
M. tenuifolia Lindl. (?M. gracilifolia Kraenzl.)
M. truncatilabia Schltr. [Near M. cobanensis Schltr. or possibly a synonym, but type is from Colombia.]
M. variabilis Batem. ex Lindl. (?M. chiriquensis Schltr., M. curtipes, Hook., M. henchmannii Hooker, M. lyonii Lindl., M. revoluta Kl., M. variabilis var. unipunctata Lindl.)

Vittariifolia Alliance

M. vittariifolia L.O.Williams

Bifrenaria rudolfii Alliance (interface with Maxillaria at M. desvauxiana?)
Bifrenaria rudolfii (Hoehne) Carnevali & G.Romero (Maxillaria rudolfii Hoehne, new name. for M. petiolaris Schltr., Maxillaria perparva Garay & Dunsterv., Bifrenaria minuta Garay, not Maxillaria minuta Cogn.)

Alliance Unknown

Camaridium boliviense Rolfe ex Stapf [description and type needed]
C. luteobrunneum Kraenzl. [description and type needed]
C. quercicola Schltr. [type needed]. Schlechter was not able to suggest a species related to this. Apparently the plant has pseudobulbs placed very distantly on a rhizome, and whitish waxy yellow flowered suffused with red with sepals about 2 cm long.

M. amblyantha Kraenzl. [description and type needed]
M. alata Ruiz & Pav., not Camaridium alatum Lindl.
M. angustibulbosa C.Schweinf. [Does this belong with Lepidota Alliance?]
M. antioquiana Kraenzl. [description and type needed]
M. barkeri Lindl. ex Beer [description and type needed]
M. binoti De Willd. [description and type needed]
M. brasiliensis Brieger & R.D.Illg. [description and drawings, etc., needed]
M. chapadensis Barb. Rodr. [listed by Cal Dodson from Kew List]
M. cinnabarina Lindl. ex J.C.Stevens, catalogue listing only.
M. connellii Rolfe [Grandiflora Alliance?]
M. convencionis Kraenzl. [description and type needed]
M. diamantensis Kraenzl. [description and type needed]
M. erubescens Kraenzl. [description and type needed]
M. galeottiana A. Rich. [Louis O. Williams (1951) could not find a drawing or type at Paris which presumably was collected in Mexico. The description indicates that the pseudobulbs are bifoliate, and that the scape is very short, features that do not correspond with any species of Maxillariinae known from Mexico.
M. graminea Lindl. [Type needed]
M. gymnochila Kraenzl. [description and type needed]
M. heterobulbon Kraenzl. [description and type needed]
M. houtteii Beer [description and type needed]
M. kreysigii Rchb.f. ex Beer [description needed], non M. kreysigii [Regel?]
M. linguaeformis Regel [Regel compares the species with M. picta, but the description indicates that the pseudobulbs are apically unifoliate]
M. luteograndiflora Hort. ex. Flor. Mag. 10: t. 599. [description needed]
M. littoralis Rouy. [description needed]
M. ornithoglossa Lodd. ex Baxt. [published in a list]
M. parthoni Dum. [description needed. Kew Index indicates "Bras." but not treated by Pabst.]
M. phaeoglossa Schltr. [description needed, illustration available similar to M. porrecta]
M. podochila Kraenzl. [description needed]
M. poeppigiana Steud. (=crocea Poepp. & Endl.)
M. polypetala Steud. Index Kewensis suggests that this is a syonym of M. platypetala Ruiz & Pav.
M. prolifera Ruiz & Pav. (obscure name)
M. sinuosa Batem. ex W.Baxt. (This may be just a name without a type)
M. spegazziniana Kraenzl. [Description and type needed]
M. stenophylla Lehm. & Kraenzl. (1899), not Rchb.f. (1854)
M. stictissimum Kraenzl.
M. tocotana Schltr. (=M. stenophylla Lehm. & Kraenzl., not Rchb.f.) [Type needed]
M. turneri Nichols. [description and type needed]
M. vestita Schltr. [possibly in Lepidota Alliance, but Schlechter compares to M. fuerstenbergiana]
M. villonacensis Dodson C. polyanthum Lehm. & Kraenzl. [type needed; not M. polyantha Barb.Rodr.]
Ornithidium heterobulbon Kraenzl. [description needed]
O. nanum Rolfe (not Maxillaria nana Hook.) [type needed]
O. olivaceum Kraenzl.

Excluded taxa. Each homotypic synonym may be a heterotypic synonym of an accepted name that is not included.

Maxillaria affinis Hort. Petrop. ex. Rchb.f., not (Poepp. & Endl.) Garay. Xylobium hyacinthina.
M. aromatica Graham ex Hooker. =Lycaste aromatica
M. atropurpurea Lodd. =Bifrenaria atropurpurea
M. aurantiaca A.Rich. & Gal. The original description indicates "scapo radicali 4-5-floro."
M. aureofulva Hook. =Bifrenaria aureofulva

M. balsamea Beer =Lycaste balsamea
M. barbata Knowles & Westc. =Bifrenaria vitellina
M. barringtoniae Hort. ex Rchb.f. =Bifrenaria harrisoniae
M. barringtoniae Lodd. =Lycaste barringtoniae
M. bicolor Ruiz & Pav. "Floribus racemosis."
M. boothii Lindl. =Nidema boothii
M. brachypus Rchb.f. =Xylobium brachypus
M. bractescens Lindl. =Xylobium bractescens
M. brasiliensis Brieger, invalid. Published without Latin description or type.
M. brevispatha Kl., Otto, & Dietr. =Lycaste candida
M. brocklehurstiana Lindl. =Houlletia brocklehurstii

M. chrysoptera Beer =Lycaste chrysoptera
M. ciliata Ruiz & Pav. =Lycaste barringtoniae
M. cochleata Beer =Lycaste cochleata
M. coelia Rchb.f. & Warsz. This is probably a xylobium.
M. colleyi Batem. ex Lindl. =Batemannia colleyi
M. concava Lindl. =Xylobium concavum
M. corrugata Lindl. This is probably a xylobium.
M. costata Lindl. =Lycaste costata
M. crinita Beer =Lycaste crinita
M. cristata Lindl. =Lycaste cristata
M. cruenta Lindl. =Lycaste cruenta
M. cuneiformis Ruiz & Pav. "Floribus racemosis."
M. cyanea Beer. =Acacallis cyanea
M. cyanocheile Hoffmanns. =Pabstia placanthera
M. cylindrobulba Regel. Probably Xylobium elongatum

M. decolor Lindl. =Xylobium palmifolium
M. deppei Lodd. =Lycaste deppei

M. elongata Lindl. & Paxt. =Xylobium elongatum

M. fimbriata Poepp. & Endl. =Lycaste aff. ciliata
M. flagellifera Lindl. ex Rchb.f. =Scuticaria steelii
M. foveata Lindl. =Xylobium foveatum
M. fulvescens Beer =Lycaste fulvescens
M. fuscata "Rchb.f." ex Gard. Chron. Error based on M. fucata Rchb.f.

M. galeata Lindl. =Gongora galeata
M. galeata Scheidw, Otto, & Dietr. =Mormolyca galeata
M. gigantea Beer, not (Lindl.) Dodson =Lycaste gigantea
M. glumibracteum Hemsl. =Epidendrum glumibracteum
M. goeringii Rchb.f. =Cymbidium goeringii
M. guttata Hort. ex. Rchb.f. =Zygopetalum xanthinum

M. harrisoniae Lindl. =Bifrenaria harrisoniae
M. hastata Ruiz & Pav. "Scapo racemosis."
M. heteroclita Poepp. & Endl. =Chaubardia heteroclita
M. heynderyexii E.Morr. =Lycaste longipetala
M. hyacinthina Rchb. =Xylobium hyacinthinum
M. hypocrita Rchb.f. =Xylobium hypocritum

M. jugosa Lindl. =Pabstia jugosa

M. laricina Kraenzl. =Pityphyllum laricinum
M. lawrencei Kraenzl. ex Index Kewensis. Invalid name based on Masdevallia lawrencei Kraenzl.
M. leiantha Beer =Lycaste deppei
M. lentiginosa Lindl. =Promenaea lentiginosa
M. leucantha Kl., Otto, & Dietr. =Lycaste leucantha
M. ligulata Ruiz & Pav. "Floribus paniculatis."
M. liliacea La Llave =Govenia liliacea
M. longifolia Lindl. =Xylobium longifolium
M. longipetala Ruiz & Pav. =Lycaste longipetala
M. luteola Beer =Polystachya luteola [=Polystachya concreta]
M. lyncea Lindl. =Stanhopea tigrina

M. macrobulbon Hook. =Lycaste macrobulbon
M. macrophylla Poee\pp. & Endl. =Lycaste macrophylla
M. maculata Lindl. Basionym: Dendrobium maculatum Kunth "Scapus...multiflorus."
M. miliacea Rchb.f. =Xylobium miliaceum

M. ornata Kl., Otto, & Dietr. =Xylobium ornatum

M. pallidiflora Hook. =Xylobium pallidiflorum
M. palmifolia Lindl. =Xylobium palmifolium
M. paniculata Ruiz & Pav. "Floribus paniculatis."
M. paraensis Brieger, invalid name: without Latin description or designation of type.
M. parvilabia Rolfe. This is a spelling error made by Rolfe based on M. parviloba Rolfe.
M. parvula Hook. =Bifrenaria parvula
M. petiolaris Schltr. =Bifrenaria rudolphii
M. phyllomega Steud. =Lycaste macrophylla
M. plaeanthera Hook. =Pabstia viridis
M. psittacina Batem. ex Steud. Name only.
M. pubigera Kl. Otto & Dietr. =Bifrenaria harrisoniae
M. purpurascens Know. & Westc. =Bifrenaria parvula
M. purpurea Beer =Polystachya purpurea

M. racemosa Hook. =Bifrenaria racemosa
M. rebellis Rchb.f. =Xylobium rebelle
M. roezlii Rchb.f. ex Lind., Name only.
M. rollinsonii Steud., orthographic error based on M. rollissonii Lindl. =Zygopetalum rollissonii
M. rollissonii Lindl. =Zygopetalum rollissonii
M. rudolfii Hoehne =Bifrenaria rudolfii

M. scabrilinguis Lindl. =Xylobium scabrilingue
M. skinneri Batem. ex Lindl. =Lycaste skinneri
M. spathacea Lindl. =Bifrenaria harrisoniae
M. squalens Hook.=Xylobium squalens
M. stachyobiora Rchb.f. =Xylobium stachyobiorum
M. stapelioides (Link & Otto) Lindl. =Promenaea stapelioides
M. steelii Hook. =Scuticaria steelii
M. stenobulbon Kl. =Xylobium pallidiflorum fide Dunsterville & Garay
M. stenopetala Know. & Westc. =Bifrenaria aureo-fulva
M. sulfurina Lem. =Xylobium sulfurinum
M. superba La Llave & Lex. =Govenia superba
M. supina Poepp. & Endl. =Xylobium supinum

M. tetragona Lindl. =Lycaste tetragona
M. tricolor Kl. Otto & Dietr. =Lycaste tricolor
M. tricolor Ruiz & Pav. "Floribus racemosis."
M. triphylla Ruiz & Pav. "Floribus racemosis."
M. truxillensis Rchb.f. = Xylobium truxillense
M. tyrianthina Hort. ex W.Baxt. =Bifrenaria tyrianthina

M. undatiflora Ruiz & pav. "Floribus paniculatis."
M. undulata Ruiz & Pav. "Scapo racemoso."

M. varicosa Rchb.f. =Xylobium varicosum
M. variegata Ruiz & Pav. =Xylobium variegatum
M. virginalis Scheidw. =Lycaste skinneri
M. viridis Lindl. =Pabstia viridis
M. vitelina lindl. =Bifrenaria vitellina

M. wageneri (Rchb.f.) Rchb.f. =Bifrenaria wageneri
M. warreana Lodd. ex Lindl. =Warrea warreana

M. xanthina Lindl. =Promenaea xanthina
M. (lyothii A.D.Hawkes, invalid name published without Latin description.

MORMOLYCA (Near Alpestris or Rufescens Alliances?)

M. aurorae D.E.Benn. & Christenson
M. galeata (Scheidw.) Garay & Wirth
M. gracilipes (Schltr.) Garay & Wirth
[Maxillaria piestopus Schltr.?]
M. polyphylla Garay & Wirth
M. schweinfurthiana Garay & Wirth
M. fuchsii J.T.Atwood (=Maxillaria?)
M. peruviana C.Schweinf.
M. ringens (Lindl.) Schltr. (=Trigonidium ringens Lindl., Mormolyca lineolata Fenzl). Hypothesis: close to Rufescens Alliance


P. amesianum Schltr.
P. antioquiense Schltr.
P. hirtzii Dodson
P. laricinum (Kraenzl.) Schltr. (=Maxillaria laricina Kraenzl.)
P. pinoides H.R.Sweet


Acuminatum Alliance [Single narrow apical leaf]

T. acuminatum Batem. ex Lindl. (T. turbinatum Rchb.f., T. subrepens Rolfe, T. tenue Lodd.)
T. pervuvianum Schltr.
T. turbinatum Rchb.f.

Egertonianum Alliance

T. egertonianum Batem. Ex Lindl. (=T. seemannii Rchb.f., ?Maxillaria brachyglossa Rich & Gal.,
T. brachyglossum (Rich. & Gal.) Schltr.), T. insigne Rchb.f. ex Benth. & Hook., T. seemannii Rchb.f., T. spathulatum Lind. & Rchb.f., T. callistele Rchb.f.
T. riopalenquense Dodson

Lankesteri Alliance

T. aurorae D.E.Benn. & Christenson
T. christensonii D.E.Benn.
T. grande Garay
T. lankesteri Ames (=T. amparoanum Schltr.)

Latifolium Alliance

T. latifolium Lindl.
T. loretoensis Schltr.
T. macranthum Barb.Rodr.
T. obtusum Lindl.

Excluded Taxa

T. amesianum, unpublished name on herbarium sheet at AMES
T. cucullatum Rchb.f., unpublished name on herbarium sheet at AMES
T. equitans Garay (=Maxillaria bicallosa (Rchb.f.) Garay)
T. monophyllum Griesb. (=Neocogniauxia monophylla (Griseb.) Schlechter)

INDEX OF TAXA [Alliances to be completed]

Aeranthus lehmannii Rchb.f.


Anthosiphon roseans Schltr.
Anthosiphon sp.

Bifrenaria minuta Garay
Bifrenaria rudolfii Alliance
Bifrenaria rudolfii (Hoehne) Carnevali & G.Romero

Broughtonia alba (Hook.f.) Spreng.

Camaridium (Barb.Rodr.) Hoehne
Camaridium adolphii Schltr.
Camaridium alatum Lindl.
Camaridium album (Hook.f.) Hoehne
Camaridium amazonicium Schltr.
Camaridium amparoana Schltr.
Camaridium arachnites Schltr.
Camaridium arbuscula Lindl.
Camaridium aureum (Poepp. & Endl.) Hoehne
Camaridium aureum var. giganteum Hoehne

Camaridium biolleyi (Schltr.) Schltr.
Camaridium boliviense Rolfe ex Stapf
Camaridium bracteatum (Schltr.) Schltr.
Camaridium bradeorum Schltr
Camaridium brenesii Schltr.
Camaridium brevifolium Lindl.

Camaridium caquetanum Schltr.
Camaridium carinatum (Barb.Rodr.) Hoehne
Camaridium cassapense (Rchb.f.) Schltr.
Camaridium caucanum Schltr.
Camaridium christobalense (Rchb.f.) Hoehne
Camaridium coccineum (Jacq.) Hoehne
Camaridium costaricense Schltr.
Camaridium cryptodanthum Authors, not Barb. Rodr.
Camaridium cryptopodanthum Authors, not Barb. Rodr.
Camaridium cyrtopodanthum Barb. Rodr.

Camaridium dendrobioides Schltr.
Camaridium densifolium (Poepp. & Endl.) Benth. & Hook.
Camaridium dichotomum Schltr.)

Camaridium equitans Schltr.
Camaridium exaltatum Kraenzl.

Camaridium flavum Schltr.

Camaridium graminifolium (Kunth) Rchb.f.)
Camaridium grandiflorum Ames, not Schltr.

Camaridium hoehnei Pabst

Camaridium iguapense (Hoehne & Schltr.) Hoehne
Camaridium imbricatum (Barb.Rodr.) Hoehne not Schltr.
Camaridium imbricatum Schltr.
Camaridium inflexum Lindl.

Camaridium jimenezii Schltr.

Camaridium lamprochlamys Schltr.
Camaridium lancifolium Rchb.f.)
Camaridium latifolium Schltr.
Camaridium lawrenceanum Rolfe
Camaridium loefgrenii (Cogn.) Hoehne
Camaridium longibracteatum Lindl.
Camaridium longum Schltr.
Camaridium luteo-brunneum Kraenzl.
Camaridium luteorubrum (Lehm. & Kraenzl.) Schltr.
Camaridium luteorubrum Lindl.
Camaridium lutescens (Scheidw.) Rchb.f.

Camaridium mathewsii (Rchb.f.) Schltr.
Camaridium minus Schltr.
Camaridium multicaule (Poepp. & Endl.) Hoehne

Camaridium nutantiflorum Schltr.

Cameridium ochroleucum Lindl.

Camaridium parviflorum Fawc.
Camaridium pendulum Barb.Rodr.
Camaridium polyanthum Lehm. & Kraenzl.
Camaridium polyphyllum (Rchb.f.) Schltr.
Camaridium purpuratum Lindl.
Camaridium purpureum Spreng.

Camaridium quercicola Schltr.

Camaridium rigidum Schltr.
Camaridium robustum Barb.Rodr.
Camaridium rugosum (Schltr.) Hoehne

Camaridium scorpioideum (Kraenzl.) Hoehne
Camaridium simile Schltr.
Camaridium sodiroi Schltr.
Camaridium squamatum (Barb. Rodr.) Hoehne
Camaridium sterrocaulon Schltr.

Camaridium tonduzii Schltr.
Camaridium tuerckheimii Schltr.

Camaridium uncatum (Lindl.) Hoehne

Camaridium vagans Schltr.
Camaridium vandiforme Schltr.
Camaridium vestitum (Sw.) Lindl.
Camaridium vinosum Rolfe, not Schltr.
Camaridium vinosum Schltr.

Camaridium wercklei Schltr.
Camaridium wrightii Schltr.

Camaridium xylobiichilum Kraenzl.

Centroglossa peruviana Cogn.

CHRYSOCYCNIS Lind. & Rchb.f.

Chrysocycnis ecuadorense Dodson & Garay
Chrysocycnis glumaceum L.O.Williams
Chrysocycnis glumaceum L.O.Williams
Chrysocycnis lehmannii Rolfe
Chrysocycnis rhomboglossa Lehm. & Kraenzl.
Chrysocycnis schlimii Linden & Rchb.f.
Chrysocycnis tigrinum (C.Schweinf.) J.T.Atwood.
Chrysocycnis trigonidii Lind. & Rchb.f.
Chrysocycnis tripterum Schltr.

Colax parkeri Spreng.


Cryptocentrum alata (Lindl.) Rausch.
Cryptocentrum brenesii, name only
Cryptocentrum caespitosum Carnevali (ined.)
Cryptocentrum calcaratum (Schltr.) Schltr.
Cryptocentrum chocoensis Hawkes
Cryptocentrum costaricensis (Schltr.), comb. ined.
Cryptocentrum dodsonii Carnevali (ined.)
Cryptocentrum dunstervilleorum Carnevali & G.A.Romero
Cryptocentrum escobarii Carnevali (ined.)
Cryptocentrum flavum Schltr.
Cryptocentrum frontinoensis (Garay) Rauschert
Cryptocentrum gracilipes Schltr.
Cryptocentrum gracillimum Ames & C.Schweinf.
Cryptocentrum gracillimum of authors, not Ames & C.Schweinf.
Cryptocentrum hirtzii Dodson
Cryptocentrum hoppii Schltr.)
Cryptocentrum inaequisepalum C.Schweinf.
Cryptocentrum jamesonii Benth.
Cryptocentrum jamesonii of authors, not Benth.
Cryptocentrum latifolium Schltr.
Cryptocentrum latifolium var. brachypetalum Garay
Cryptocentrum lehmannii (Rchb.f.) Garay
Cryptocentrum longipetalum Carnevali (ined.)
Cryptocentrum longiscapum Brieger
Cryptocentrum minus Schltr.
Cryptocentrum pergracile Schltr.
Cryptocentrum peruvianum (Cogn.) C.Schweinf. ssp. peruvianum
Cryptocentrum peruvianum ssp. dactylinum Carnevali, ined.
Cryptocentrum peruvianum ssp. minus (Schltr.) Carnevali, ined.
Cryptocentrum pervuianum of authors, not (Cogn.) C.Schweinf.
Cryptocentrum pseudobulbosum C.Schweinf.
Cryptocentrum rhomboglossa (Lehm. & Kraenzl.) Rauschert
Cryptocentrum roseans (Schltr.) A.D.Hawkes
Cryptocentrum silverstonei Carnevali (ined.)
Cryptocentrum spathaceum Dodson
Cryptocentrum standleyii Ames
Cryptocentrum strumpflei (Garay) Rauschert
Cryptocentrum subgenus Cryptocentrum
Cryptocentrum subgenus Polysticha Carnevali
Cryptocentrum subgenus Pseudobulbosa Carnevali

Cymbidium ochroleucum Lindl.
Cymbidium proliferum (Sw.) Sw.
Cymbidium vestitum (Sw.) Sw.


Cyrtidiorchis alatum (Lindl.) Rauschert
Cyrtidiorchis frontinoensis (Garay) Rausch.
Cyrtidiorchis rhomboglossum (Lehm. & Kraenzl.) Rauschert
Cyrtidiorchis stumpflei (Garay) Rausch.
Cyrtidiorchis triptera (Schltr.) Rausch.

Cyrtidium Schltr.
Cyrtidium alatum (Lindl.) Garay
Cyrtidium frontinoensis Garay
Cyrtidium rhomboglossum (Lehm. & Kraenzl.) Schltr.
Cyrtidium stumpflei Garay
Cyrtidium tripterum (Schltr.) Schltr.

Dendrobium album Hook.f.
Dendrobium latifolium Kunth.
Dendrobium maculatum Kunth.
Dendrobium platypetalum (Ruiz & Pav.) Pers.
Dendrobium ramosum Pers.

Dicrypta (Heterotaxis) Alliance
Dicrypta baueri Lindl.
Dicrypta crassifolia (Lindl.) Lindl. ex Loud.
Dicrypta densifolia Poepp. & Endl.
Dicrypta discolor Lodd. ex Lindl.
Dicrypta elatior Rchb.f.
Dicrypta iridifolia Bateman ex Rchb.f.
Dicrypta irisphyta Barb.Rodr.
Dicrypta villosa Barb. Rodr.

Epidendrum coccineum Jacq.
Epidendrum proliferum Sw.
Epidendrum sessile Sw.
Epidendrum vestitum Sw.

Fernandezia haemathodes Ruiz & Pav.
Fernandezia punctata Ruiz & Pav.

Heterotaxis crassifolia Lindl.

Isochilus prolifer (Sw.) R.Br.
Isochilus graminifolius Kunth
Isochilus grandiflorum Lindl.

Kefersteinia bicallosa (Rchb.f.) Rchb.f.

Lycaste aciantha (Rchb.f.) Rchb.f.

Marsupiaria iridifolia Hoehne
Marsupiaria valenzuelana (A.Rich) Garay
Marsupiaria vandiformis (Schltr.) Hoehne


Maxillaria X dunstervillei Carnevali & Ramirez
Maxillaria X lyothii A.D.Hawkes)?
Maxillaria abelei Schltr.
Maxillaria acervata Rchb.f.
Maxillaria aciantha Rchb.f.
Maxillaria acicularis Herb.
Maxillaria acostaei Schltr.
Maxillaria Acuminata Alliance
Maxillaria acuminata Lindl.
Maxillaria acutifolia Lindl., not Schltr.
Maxillaria acutifolia Schltr., not Lindl.
Maxillaria acutipetala Hook.
Maxillaria adendrobium (Rchb.f.) Dressler
Maxillaria admonens Bock
Maxillaria adolphii Ames & Correll
Maxillaria adscendens Schltr.
Maxillaria aequiloba Schltr.
Maxillaria affinis (Poepp. & Endl.) Garay
Maxillaria affinis Hort. Petrop. ex. Rchb.f., not (Poepp. & Endl.) Garay.
Maxillaria aggregata (H.B.K.) Lindl.
Maxillaria alata Ruiz & Pav.
Maxillaria Alba Alliance
Maxillaria alba (J.D.Hook.) Lindl.
Maxillaria Albata Alliance
Maxillaria albata Lindl.
Maxillaria albertii Schltr.
Maxillaria albiflora Ames & C.Schweinf.
Maxillaria alfaroi Ames & C.Schweinf.
Maxillaria allenii L.O.Williams
Maxillaria Alpestris Alliance
Maxillaria alpestris Lindl.
Maxillaria alticola C.Schweinf.
Maxillaria Amabilis Suballiance
Maxillaria amabilis J.T.Atwood
Maxillaria amazonica
Maxillaria amblyantha Kraenzl.
Maxillaria amesiana Hort. ex Gard. Chron.
Maxillaria amparoana Schltr.
Maxillaria ampliflora C.Schweinf.
Maxillaria anatomorum Rchb.f.
Maxillaria anceps Ames & C.Schweinf.
Maxillaria angustibulbosa C.Schweinf.
Maxillaria angustifolia Hook.
Maxillaria angustisegmenta Ames, Hubbard, & C.Schweinf.
Maxillaria angustissima Ames, Hubbard, & C.Schweinf.
Maxillaria antioquiana Kraenzl.
Maxillaria appendiculoides C.Schweinf.
Maxillaria arachnites Rchb.f.
Maxillaria arachnitiflora Ames & Correll
Maxillaria Arbuscula Suballiance
Maxillaria arbuscula (Lindl.) comb. ined., new name perhaps needed
Maxillaria arbuscula Rchb.f.
Maxillaria argyrophylla Poepp. & Endl.
Maxillaria aristeguietae Foldats
Maxillaria aromatica Graham ex Hooker.
Maxillaria articulada Kl.
Maxillaria atrata Rchb.f.
Maxillaria atrata var. brachyantha Rchb.f.
Maxillaria atropurpurea Lodd.
Maxillaria attenuata Ames & C.Schweinf.
Maxillaria augustaevictoriae Lehm. & Kraenzl. ex Kraenzl.
Maxillaria aurantiaca A.Rich. & Gal.
Maxillaria aurantiaca Schltr., not A.Rich. & Gal.
Maxillaria Aurea Suballiance
Maxillaria aurea (Poepp. & Endl.) L.O.Williams
Maxillaria aurea Hort. ex Rchb.f.
Maxillaria aureofulva Hook.
Maxillaria aurorae D.E.Benn. & Christenson
Maxillaria auyantepuiensis Foldats
Maxillaria auyantepuiensis ssp. epiphytica Carnevali & Ramirez

Maxillaria balsamea Beer
Maxillaria barbata Knowles & Westc.
Maxillaria barbosae Loefgren ex Porto
Maxillaria barkeri Lindl. ex Beer
Maxillaria barringtoniae Hort. ex Rchb.f.
Maxillaria barringtoniae Lodd.
Maxillaria batemanii Poepp. & Endl.
Maxillaria baumanniana Schltr.
Maxillaria bennettii Christenson
Maxillaria Bicallosa Suballiance
Maxillaria bicallosa (Rchb.f.) Garay
Maxillaria bicolor (Rolfe) ined.
Maxillaria bicolor Ruiz & Pav.
Maxillaria binoti De Willd.
Maxillaria Biolleyi Alliance
Maxillaria Biolleyi Suballiance
Maxillaria biolleyi (Schltr.) L.O.Williams
Maxillaria bolivarensis C.Schweinf.
Maxillaria boliviensis Schltr.
Maxillaria bolleoides Schltr.
Maxillaria bomboizensis Dodson
Maxillaria boothii Lindl.
Maxillaria brachybulbon Schltr.
Maxillaria brachyglossa Rich & Gal.
Maxillaria brachypetala Schltr.
Maxillaria brachypoda Schltr.
Maxillaria brachypus Rchb.f.
Maxillaria bracteata (Schltr.) Ames & Correll
Maxillaria bractescens Lindl.
Maxillaria bradei Schltr. ex Hoehne
Maxillaria Bradeorum Suballiance
Maxillaria bradeorum (Schltr.) L.O.Williams
Maxillaria brasiliensis Brieger, invalid
Maxillaria brasiliensis Brieger & Bicalho
Maxillaria brasiliensis Brieger & R.D.Illg.
Maxillaria brenesii Schltr.
Maxillaria brenesii var. longiloba (Ames & C.Schweinf.) J.T.Atwood
Maxillaria brevifolia (Lindl.) Rchb.f.
Maxillaria brevilabia Ames & Correll
Maxillaria brevipedunculata Ames & C.Schweinf.
Maxillaria brevipes Schltr.
Maxillaria brevis (Hoehne & Schltr.) Hoehne
Maxillaria breviscapa Poepp. & Endl.
Maxillaria brevispatha Kl., Otto, & Dietr.
Maxillaria broadwayi (Cogn.) R.E.Schultes
Maxillaria brocklehurstiana Lindl.
Maxillaria brunnea Lind. & Rchb.f.
Maxillaria buchtienii Schltr.
Maxillaria burgeri J.T.Atwood

Maxillaria cacaoensis J.T.Atwood
Maxillaria caespitifica Rchb.f.
Maxillaria caespitosa C.Schweinf.
Maxillaria calantha Schltr.
Maxillaria callichroma Rchb.f.
Maxillaria caloglossa Rchb.f.
Maxillaria Camaridii Alliance
Maxillaria Camaridii Suballiance
Maxillaria camaridii Rchb.f.
Maxillaria camaridioides Schltr.
Maxillaria campanulata C.Schweinf.
Maxillaria candida Lodd. ex Lindl.
Maxillaria caparaoensis Brade
Maxillaria caquetana Schltr.
Maxillaria carinata Barb. Rodr.
Maxillaria carinulata Rchb.f.
Maxillaria cassapensis Rchb.f.
Maxillaria casta Kraenzl.
Maxillaria caucae Garay
Maxillaria caucana Schltr.
Maxillaria caucana (Schltr.) comb. ined.
Maxillaria caulina Schltr.
Maxillaria caveroi D.E.Benn. & Christenson
Maxillaria cedralensis J.T.Atwood & D.E.Mora
Maxillaria cepula Rchb.f.
Maxillaria cerifera Barb.Rodr.
Maxillaria chacoensis Dodson
Maxillaria chapadensis Barb. Rodr.
Maxillaria chartacifolia Ames & C.Schweinf.
Maxillaria chasei Dodson
Maxillaria Chicana Alliance
Maxillaria chicana Dodson
Maxillaria chionantha J.T.Atwood
Maxillaria chiriquensis Schltr.
Maxillaria chlorantha Lindl.
Maxillaria chlorochila Lehm. & Kraenzl. ex Kraenzl.
Maxillaria christobalensis Rchb.f.
Maxillaria chrysantha Barb.Rodr.
Maxillaria chrysocycnoides (Schltr.) Dodson
Maxillaria chrysoptera Beer
Maxillaria ciliata Ruiz & Pav.
Maxillaria cinnabarina Lindl. ex J.C.Stevens
Maxillaria cleistogama Brieger & Bicalho
Maxillaria cobanensis Schltr.
Maxillaria Coccinea Suballiance
Maxillaria coccinea (Jacq.) L.O.Williams ex Hodge
Maxillaria cochleata Beer
Maxillaria coelia Rchb.f. & Warsz.
Maxillaria cogniauxiana Hoehne
Maxillaria colleyi Batem. ex Lindl.
Maxillaria colorata Rchb.f.
Maxillaria compacta (Schltr.) comb. ined.
Maxillaria compressibulba Schltr.
Maxillaria concava Lindl.
Maxillaria concavilabia Ames & Correll
Maxillaria condensata C.Schweinf.
Maxillaria conduplicata (Ames & C.Schweinf.) L.O.Williams
Maxillaria conferta Ames & C.Schweinf.
Maxillaria confusa Ames & C.Schweinf.
Maxillaria connellii Rolfe
Maxillaria consanguinea Kl.
Maxillaria convencionis Kraenzl.
Maxillaria cordyline (Rchb.f.) Dodson
Maxillaria coriacea Barb. Rodr.
Maxillaria cornuta C.Schweinf.
Maxillaria corrugata Lindl.
Maxillaria costaricensis Schltr.
Maxillaria costata Lindl.
Maxillaria cozieriana H.G.Jones
Maxillaria crassicaulis C.Schweinf.
Maxillaria crassicaulis C.Schweinf.
Maxillaria Crassifolia Suballiance
Maxillaria crassifolia (Lindl.) Rchb.f.
Maxillaria crassipes Kraenzl.
Maxillaria crinita Beer
Maxillaria cristata Lindl.
Maxillaria crocea Lindl. not Poepp. & Endl.
Maxillaria crocea Poepp. & Endl.
Maxillaria croceorubens (Rchb.f.) L.O.Williams
Maxillaria cruenta Lindl.
Maxillaria cryptobulbon Carnevali & J.T.Atwood
Maxillaria Ctenostachya Alliance
Maxillaria ctenostachya Rchb.f.
Maxillaria Cucullata Alliance
Maxillaria cucullata Lindley
Maxillaria cuencana Garay
Maxillaria cuneiformis Ruiz & Pav.
Maxillaria curtipes, Hook.
Maxillaria cuzcoensis C.Schweinf.
Maxillaria cyanea Beer.
Maxillaria cyanocheile Hoffmanns.
Maxillaria cylindrobulba Regel.
Maxillaria cymbidioides Dodson, J.T.Atwood, & Carnevali

Maxillaria dalessandroi Dodson
Maxillaria dalstromii Dodson
Maxillaria darienensis J.T.Atwood
Maxillaria decolor Lindl.
Maxillaria deflexa Kl. ex A.Br.Kl. & Bouche
Maxillaria dendrobioides (Schltr.) L.O.Williams
Maxillaria Densa Alliance
Maxillaria densa Lindl.
Maxillaria Densifolia Alliance
Maxillaria Densifolia Suballiance
Maxillaria densifolia (Poepp. & Endl.) Rchb.f.
Maxillaria deppei Lodd.
Maxillaria Desvauxiana Alliance
Maxillaria desvauxiana Rchb.f.
Maxillaria diamantensis Kraenzl.
Maxillaria dichotoma (Schltr.) L.O.Williams
Maxillaria dichroma Rolfe
Maxillaria dillonii D.E.Benn. & Christenson
Maxillaria discolor (Lodd. ex Lindl.) Rchb.f.
Maxillaria disticha (Lindl.) C.Schweinf.
Maxillaria diuturna Ames & C.Schweinf.
Maxillaria divaricata (Barb.Rodr.) Cogn.
Maxillaria divitiflora Rchb.f.
Maxillaria dodsonii Carnevali, ined.
Maxillaria dolichophylla Schltr.
Maxillaria dressleriana Carnevali & J.T.Atwood

Maxillaria eburnea Lindl.
Maxillaria echinophyta Barb.Rodr.
Maxillaria echinochila Kraenzlin
Maxillaria ecuadorensis Schltr.
Maxillaria edwardsii D.E.Benn. & Christenson
Maxillaria elata Schltr.
Maxillaria Elatior Alliance
Maxillaria elatior (Rchb.f.) Rchb.f.
Maxillaria elegans Schltr.
Maxillaria elegantula Rolfe
Maxillaria elongata Lindl. & Paxt.
Maxillaria embreei Dodson
Maxillaria encyclioides Dodson & J.T.Atwood
Maxillaria endresii Rchb.f.
Maxillaria endresii Rchb.f. var. angustisegmenta (Ames, Hubbard, & C.Schweinf.) C. Schweinf.
Maxillaria equitans (Schltr.) Garay
Maxillaria erubescens Kraenzl.
Maxillaria estradae Dodson
Maxillaria exaltata (Kraenzl.) C. Schweinf.
Maxillaria exigua Regel

Maxillaria falcata Ames & C.Schweinf.
Maxillaria fallax Schltr.
Maxillaria farinifera Schltr.
Maxillaria fasciculata C.Schweinf.
Maxillaria ferdinandiana Barb.Rodr
Maxillaria fimbriata Poepp. & Endl.
Maxillaria flagellifera Lindl. ex Rchb.f.
Maxillaria flava Ames, Hubbard, & C.Schweinf.
Maxillaria flavoviridis Barb.Rodr.
Maxillaria flavum (Schltr.) comb. ined.
Maxillaria fletcheriana Rolfe
Maxillaria Floribunda Alliance
Maxillaria floribunda Lindl.
Maxillaria foldatsiana Carnevali & Ramirez
Maxillaria foliosa Ames & C.Schweinf.
Maxillaria foveata Lindl.
Maxillaria fractiflexa Rchb.f.
Maxillaria Friedrichsthalii Alliance
Maxillaria friedrichsthalii Rchb.f.
Maxillaria fritzii Carnevali & Dodson, ined.
Maxillaria fucata Rchb.f.
Maxillaria fuerstenbergiana Schltr.
Maxillaria fulgens (Rchb.f.) L.O.Williams
Maxillaria fulvescens Beer
Maxillaria funerea Lindl.
Maxillaria funicaulis C.Schweinf.
Maxillaria furfuracea Scheidw.
Maxillaria fuscata "Rchb.f." ex Gard. Chron.
Maxillaria fuscata Hort. ex Rchb.f. 1863, non Kl. ex Rchb.f.
Maxillaria fuscata Kl. ex Rchb.f. [1863], non Hort. ex Rchb.f. [1863]
Maxillaria fuscopurpurea Drapiez

Maxillaria galantha J.T.Atwood & Carnevali
Maxillaria galeata Lindl.
Maxillaria galeata Scheidw, Otto, & Dietr.
Maxillaria galeottiana A. Rich.
Maxillaria galeottiana Regel, not Rich. et Gal.
Maxillaria gatunensis Schltr.
Maxillaria gentryi Dodson
Maxillaria gigantea (Lindl.) Dodson
Maxillaria gigantea Beer, not (Lindl.) Dodson
Maxillaria gigantea ssp. condorensis D.E.Benn. & Christenson
Maxillaria glauca Lindl. ex W. Baxt.
Maxillaria glomerata Gal.
Maxillaria glumacea Rolfe
Maxillaria glumibracteum Hemsl.
Maxillaria goeringii Rchb.f.
Maxillaria gomeziana J.T.Atwood
Maxillaria gracilifolia Kraenzl.
Maxillaria gracilipes Schltr.
Maxillaria gracilis Lodd.
Maxillaria graminea Lindl.
Maxillaria Graminifolia Alliance
Maxillaria graminifolia (Kunth) Rchb.f.
Maxillaria Grandiflora Alliance
Maxillaria grandiflora (HBK) Lindl.
Maxillaria grandimentum C.Schweinf.
Maxillaria grandis Rchb.f.
Maxillaria granditenuis D.E.Benn. & Christenson
Maxillaria grayi Dodson
Maxillaria Grobyoides Suballiance
Maxillaria grobyoides Garay & Dunsterville
Maxillaria guadalupensis Cogn.
Maxillaria guareimensis Rchb.f.
Maxillaria guayanensis Klinge
Maxillaria guentheriana Kraenzl.
Maxillaria guiardiana Chiron
Maxillaria guttata Hort. ex. Rchb.f.
Maxillaria gymnochila Kraenzl.

Maxillaria haberi J.T.Atwood
Maxillaria haemathodes (Ruiz & Pav.) Garay
Maxillaria haenkei Correll
Maxillaria hagsateriana Soto Arenas
Maxillaria harrisoniae Lindl.
Maxillaria hastata Ruiz & Pav.
Maxillaria Hastulata/Pulla Suballiance
Maxillaria hastulata Lindl.
Maxillaria hatschbachii (Schltr.) Hoehne
Maxillaria hedwigiae Hamer & Dodson
Maxillaria hedyosma Schltr.
Maxillaria heidelbergensis Senghas, illeg.
Maxillaria heismanniana Barb.Rodr.
Maxillaria hematoglossa Rich. & Gal.
Maxillaria henchmannii Hooker
Maxillaria hennisiana Schltr.
Maxillaria heraldica Rchb.f.
Maxillaria herzogiana Kraenzl.
Maxillaria heterobulbon Kraenzl.
Maxillaria heteroclita Poepp. & Endl.
Maxillaria heterophylla Hoehne
Maxillaria heynderyexii E.Morr.
Maxillaria hillsii Dodson
Maxillaria hirsutilabia D.E.Benn. & Christenson
Maxillaria hirtilabia Lindl.
Maxillaria hirtzii Dodson & A.Andreetta
Maxillaria histiologorum Rchb.f., fide Pabst
Maxillaria histrionica (Rchb.f.) L.O.Williams
Maxillaria hoehneana P.F.Hunt
Maxillaria hoehnei Schltr.
Maxillaria hoppii Schltr.
Maxillaria horichii Senghas
Maxillaria houtteana Rchb.f.
Maxillaria houtteii Beer
Maxillaria huancabambae (Kraenzl.) C.Schweinf.
Maxillaria huancabambae (Kraenzl.) C.Schweinf. var. cuzcoensis C.Schweinf.
Maxillaria huebneri Schltr.
Maxillaria hyacinthina Rchb.
Maxillaria hypocrita Rchb.f.

Maxillaria iguapensis Hoehne & Schltr.
Maxillaria Imbricata Alliance
Maxillaria imbricata Barb.Rodr.
Maxillaria Inaudita Suballiance
Maxillaria inaudita Rchb.f.
Maxillaria inaudita var. minor Ames & C.Schweinf.
Maxillaria infausta Rchb.f.
Maxillaria inflexa (Lindl.) Griseb.
Maxillaria insignis Rolfe
Maxillaria insolita Dressler
Maxillaria iridifolia (Batem. ex Rchb.f.) Rchb.f.
Maxillaria irrorata Rchb.f.

Maxillaria jamboensis Dodson
Maxillaria jamesonii (Rchb.f.) Garay & C.Schweinf.
Maxillaria jenischiana Rchb.f.
Maxillaria johannis Pabst
Maxillaria johniana Kraenzl.
Maxillaria jucunda Lehm. & Kraenzl.
Maxillaria juergensii Schltr.
Maxillaria jugata Garay
Maxillaria jugosa Lindl.

Maxillaria kalbreyeri Rchb.f.
Maxillaria kautskii Pabst
Maxillaria kegelii Rchb.f.
Maxillaria klugii C.Schweinf.
Maxillaria koehleri Schltr.
Maxillaria kreysigii [Regel?]
Maxillaria kreysigii Rchb.f. ex Beer, non M. kreysigii [Regel]

Maxillaria lactea Schltr. 1923 not 1925
Maxillaria lactea Schltr. 1925, not 1923
Maxillaria lactiflora Pabst
Maxillaria laevilabris Lindl.
Maxillaria laevilabris Lindl.
Maxillaria lamprochlamys (Schltr.) comb. ined.
Maxillaria lancifolia (Rchb.f.) Rchb.f.
Maxillaria langlassei Schltr.
Maxillaria lankesteri Ames
Maxillaria laricina Kraenzl.
Maxillaria lasallei Fold.
Maxillaria latifolia (Kunth) Lindl.
Maxillaria lawrenceana (Rolfe) Garay & Dunsterv.
Maxillaria lawrencei Kraenzl. ex Index Kewensis
Maxillaria lehmannii Rchb.f.
Maxillaria lehmannii (Kraenzl.) comb. ined.
Maxillaria leiantha Beer
Maxillaria lentiginosa Lindl.
Maxillaria Lepidota Alliance
Maxillaria Lepidota Suballiance
Maxillaria lepidota Lindl.
Maxillaria leptosepala Hook.
Maxillaria leucaimata Barb.Rodr.
Maxillaria leucantha Kl., Otto, & Dietr.
Maxillaria leucantha Schltr. (1929), not Kl., Otto, & Dietr. (1850)
Maxillaria leucocheile Hoffmsgg.
Maxillaria lexarzana Soto Arenas & F.Chiang
Maxillaria ligulata Ruiz & Pav.
Maxillaria liliacea Barb. Rodr. not Ll. & Lex.
Maxillaria liliacea La Llave
Maxillaria lilliputana D.E.Benn. & Christenson
Maxillaria lindeniana Rich. & Gal.
Maxillaria lindenii Cogn.
Maxillaria lindleyana Schltr.
Maxillaria linearifolia Ames & C.Schweinf.
Maxillaria linearis C.Schweinf.
Maxillaria linguaeformis Regel
Maxillaria liparophylla Summerhayes
Maxillaria litensis Dodson
Maxillaria littoralis Rouy.
Maxillaria loefgrenii (Cogn.) Pabst
Maxillaria longa (Schltr.) Dodson
Maxillaria longibracteata (Lindl.) Rchb.f.
Maxillaria longibracteata (Lindl.) Rchb.f. var. grandiflora (Lindl.) C.Schweinf.
Maxillaria longicaulis Schltr.
Maxillaria longifolia (Barb.Rodr.) Cogn. not Lindl.
Maxillaria longifolia Lindl.
Maxillaria longiloba (Ames & C.Schweinf.) J.T.Atwood
Maxillaria longipes Lindl.
Maxillaria longipetala Barb. Rodr., not Ruiz & Pav.
Maxillaria longipetala Ruiz & Pav.
Maxillaria longipetiolata Ames. & C.Schweinf.
Maxillaria longisepala Rolfe
Maxillaria longissima Lindl.
Maxillaria loretoensis C.Schweinf.
Maxillaria lorifolia Rchb.f.
Maxillaria Lueri Alliance
Maxillaria lueri Dodson
Maxillaria luteograndiflora Hort. ex. Flor. Mag. 10: t. 599.
Maxillaria luteoalba Lindl.
Maxillaria luteola Beer
Maxillaria luteorubra (Lindl.) Rchb.f.
Maxillaria lutescens Scheidw.
Maxillaria lutheri J.T.Atwood
Maxillaria lyncea Lindl.
Maxillaria lyonii Lindl.

Maxillaria machinazensis D.E.Benn. & Christenson
Maxillaria macleei Batem. ex Lindl.
Maxillaria macrobulbon Hook.
Maxillaria macrophylla Poee\pp. & Endl.
Maxillaria macropoda Schltr.
Maxillaria macrura Rchb.f.
Maxillaria maculata Lindl.
Maxillaria maderoi Schltr.
Maxillaria Madida Suballiance
Maxillaria madida Lindl.
Maxillaria maleolens Schltr.
Maxillaria mapiriensis (Kraenzl.) L.O.Williams
Maxillaria Marginata Suballiance
Maxillaria marginata Fenzl.
Maxillaria mariaisabelae J.T.Atwood
Maxillaria marmoliana Dodson
Maxillaria mathewsii Lindl., not Rchb.f.
Maxillaria mathewsii of authors, not Lindl. (1845) nor Rchb.f. (1863)
Maxillaria mathewsii Rchb.f. not Lindl.
Maxillaria mattogrossensis Brade
Maxillaria meirax Rchb.f. & Warm.
Maxillaria meleagris Lindl.
Maxillaria melina Lindl.
Maxillaria merana Dodson
Maxillaria Meridensis Alliance
Maxillaria meridensis Lindl.
Maxillaria mexicana J.T.Atwood & Soto, ined.
Maxillaria microblephara Schltr.
Maxillaria microdendron Schltr.
Maxillaria Microphyton Alliance
Maxillaria microphyton Schltr.
Maxillaria microtricha Schltr.
Maxillaria miliacea Rchb.f.
Maxillaria miniata (Lindl.) L.O.Williams
Maxillaria minor (Schltr.) L.O.Williams
Maxillaria minuta Cogn.
Maxillaria mirabilis Cogn.
Maxillaria modesta Brade not Schltr.
Maxillaria modesta Schltr. not Brade
Maxillaria modestiflora Pabst
Maxillaria molitor Rchb.f.
Maxillaria mombachoensis Heller ex J.T.Atwood
Maxillaria monacensis Kraenzl.
Maxillaria monantha Barb.Rodr.
Maxillaria monoceros Kl. ex Rchb.f
Maxillaria monteverdensis J.T.Atwood & G. Barboza
Maxillaria mooreana Rolfe
Maxillaria moralesii Carnevali & J.T.Atwood
Maxillaria mosenii Kraenzl.
Maxillaria moutinhoi ex Index Kewensis
Maxillaria moutini Pabst
Maxillaria muelleri Regel
Maxillaria multicaulis (Poepp. & Endl.) C.Schweinf.
Maxillaria multiflora Barb.Rodr.
Maxillaria murilliana Hoehne
Maxillaria muscicola Rchb.f.
Maxillaria muscoides J.T.Atwood
Maxillaria mutabilis Hort. ex Rchb.f.

Maxillaria nagelii L.O.Williams ex Correll
Maxillaria nana Hook.
Maxillaria nanegalensis Rchb.f.
Maxillaria napoensis Dodson
Maxillaria Nardoides Alliance
Maxillaria nardoides Kraenzl.
Maxillaria nasalis Rchb.f.
Maxillaria nasuta Rchb.f.
Maxillaria neglecta (Schltr.) L.O.Williams
Maxillaria neillii Dodson
Maxillaria neophylla Rchb.f.
Maxillaria neowiedii Rchb.f.
Maxillaria nervosa Rolfe ex Britton
Maxillaria nicaraguensis (Hamer & Garay) J.T.Atwood
Maxillaria nigrescens Lindl.
Maxillaria nitidula Rchb.f.
Maxillaria nivea (Lindl.) L.O.Williams
Maxillaria notylioglossa Rchb.f.
Maxillaria nubigena (Rchb.f.) C.Schweinf.
Maxillaria nuriensis Carnevali & Ramirez
Maxillaria nutans Lindl.
Maxillaria nutantiflora Schltr.
Maxillaria nymphopolitana Kraenzl.

Maxillaria obscura Lind. & Rchb.f.
Maxillaria ochracea (Rchb.f.) Garay
Maxillaria ochroglossa Schltr.
Maxillaria ochroleuca Lodd. ex Lindl.
Maxillaria ochroleuca var. longipes Sander
Maxillaria oestlundiana L.O.Williams
Maxillaria ophiodens J.T.Atwood
Maxillaria oreocharis Schltr.
Maxillaria ornata Kl., Otto, & Dietr.
Maxillaria ornithoglossa Lodd. ex Baxt.
Maxillaria oxysepala Schltr.

Maxillaria pachyacron Schltr.
Maxillaria pachyneura Lehm. & Kraenzl.
Maxillaria pachyphylla Schltr. ex Hoehne
Maxillaria Paleata (wrightii) Alliance
Maxillaria paleata (Rchb.f.) Ames & Correll
Maxillaria pallidiflava (Schltr.) Senghas
Maxillaria pallidiflora Hook.
Maxillaria pallidiflora Rchb.f. ex Barb. Rodr., illeg.
Maxillaria palmensis Dodson
Maxillaria palmifolia Lindl.
Maxillaria pamplonensis Lind. & Rchb.f.
Maxillaria panamensis Schltr.
Maxillaria paniculata Ruiz & Pav.
Maxillaria pannieri Fold.
Maxillaria pantherina Rchb.f. not Hoffmsgg.
Maxillaria paraensis Brieger
Maxillaria parahybunensis Cogn.
Maxillaria paranaensis Barb. Rodr.
Maxillaria pardalina Garay
Maxillaria Parkeri Alliance
Maxillaria parkeri (Spreng.) Hook.
Maxillaria parthoni Dum.
Maxillaria parva Rolfe
Maxillaria parvibulbosa C.Schweinf.
Maxillaria parviflora (Poepp. & Endl.) Garay
Maxillaria parvilabia Ames & C.Schweinf.
Maxillaria parvilabia Rolfe
Maxillaria parviloba Rolfe
Maxillaria parvula Hook. not Schltr.
Maxillaria parvula Schltr. not Hook.
Maxillaria pastensis Rchb.f.
Maxillaria pastoensis (Schltr.) comb. ined.
Maxillaria patens Schltr.
Maxillaria patula C.Schweinf.
Maxillaria pauciflora Barb.Rodr.
Maxillaria paulistana Hoehne
Maxillaria pendens Pabst
Maxillaria pendula (Poepp. & Endl.) C.Schweinf.
Maxillaria penduliflora Fenzl.
Maxillaria pentura Lindl.
Maxillaria perijaensis Carnevali & Romero, ined.
Maxillaria perparva Garay & Dunsterv.
Maxillaria perpusilla Kraenzl.
Maxillaria perryae Dodson
Maxillaria pertusa Lindl. ex Rchb.f.
Maxillaria petiolaris A. Rich. ex Rchb.f.
Maxillaria petiolaris Schltr.
Maxillaria peruviana (C.Schweinf.) D.E.Benn. & Christenson
Maxillaria pfitzerianum (Lehm. & Kraenzl.) comb. ined.
Maxillaria phaeoglossa Schltr.
Maxillaria phoenicanthera Barb.Rodr.
Maxillaria phyllomega Steud.
Maxillaria Picta Alliance
Maxillaria picta Hook.
Maxillaria picta Hook. var. brunnea Rchb.f.
Maxillaria picta Rchb.f. non Hook.
Maxillaria picta Hook. var. brevis Hoehne & Schltr.
Maxillaria piestopus Schltr.
Maxillaria piresiana Hoehne
Maxillaria pittieri (Ames) L.O.Williams
Maxillaria plaeanthera Hook.
Maxillaria Planicola Alliance
Maxillaria planicola C.Schweinf.
Maxillaria platyloba Schltr.
Maxillaria platypetala Ruiz & Pav.
Maxillaria plebeja Rchb.f.
Maxillaria pleiantha Schltr.
Maxillaria pleuranthioides (Schltr.) Garay
Maxillaria plicata Schltr.
Maxillaria poaefolia Schltr.
Maxillaria podochila Kraenzl.
Maxillaria poeppigiana Steud.
Maxillaria poicilothece Schltr.
Maxillaria polyantha Barb.Rodr.
Maxillaria polybulbon Kraenzl.
Maxillaria polypetala Steud.
Maxillaria polyphylla Rchb.f.
Maxillaria ponerantha of authors, not Rchb.f.
Maxillaria ponerantha Rchb.f.
Maxillaria Ponerantha Suballiance
Maxillaria porphyrostele Rchb.f.
Maxillaria Porrecta (Brunnea) Suballiance
Maxillaria porrecta Lindl.
Maxillaria powellii Schltr.
Maxillaria praestans Rchb.f.
Maxillaria praetexta Rchb.f.
Maxillaria proboscidea Rchb.f.
Maxillaria procurrens Lindl.
Maxillaria prolifera Ruiz & Pav.
Maxillaria pseudoneglecta J.T.Atwood
Maxillaria pseudoreichenheimiana Dodson
Maxillaria psittacina Batem. ex Steud
Maxillaria pterocarpa Barb.Rodr.
Maxillaria Pterocarpa Suballiance
Maxillaria pubigera Kl. Otto & Dietr.
Maxillaria pubilabia Schltr.
Maxillaria pulchra (Schltr.) L.O.Williams ex Correll
Maxillaria pulla Lind. & Rchb.f.
Maxillaria Pumila Alliance
Maxillaria pumila Hook.
Maxillaria punctata Hoehne non Lodd.
Maxillaria punctata Lodd.
Maxillaria punctata Schltr. not Lodd.
Maxillaria punctostriata Rchb.f.
Maxillaria punctulata Kl
Maxillaria purpurascens Know. & Westc.
Maxillaria purpurata (Lindl.) Rchb.f.
Maxillaria purpurea Ames & Correll
Maxillaria purpurea Beer
Maxillaria pusilla Rolfe

Maxillaria quadrata Ames & Correll
Maxillaria queirogana Barb.Rodr.
Maxillaria quelchii Rolfe
Maxillaria quitensis (Rchb.f.) C. Schweinf.

Maxillaria racemosa Hook.
Maxillaria ramonensis Schltr.
Maxillaria Ramosa Alliance
Maxillaria ramosa Ruiz & Pav.
Maxillaria ramosissima Kraenzl.
Maxillaria rebellis Rchb.f.
Maxillaria regeliana Cogn.
Maxillaria reichenheimiana Endr. & Rchb.f.
Maxillaria repens L.O.Williams
Maxillaria revoluta Kl.
Maxillaria rhodosticta Kraenzl.
Maxillaria rhombea Lindl.
Maxillaria richii Dodson
Maxillaria rigida Barb. Rodr.
Maxillaria Ringens Suballiance
Maxillaria ringens Rchb.f.
Maxillaria riopalenquensis Dodson
Maxillaria robusta Barb.Rodr.
Maxillaria rodriguesii Cogn.
Maxillaria rodrigueziana J.T.Atwood & Mora
Maxillaria roezlii Rchb.f. ex Lind.
Maxillaria rollinsonii Steud.
Maxillaria rollissonii Lindl.
Maxillaria rotundilabia C.Schweinf.
Maxillaria rouseauae Schltr.
Maxillaria ruberrima (Lindl.) Garay
Maxillaria rubioi Dodson
Maxillaria rubrilabia Schltr.
Maxillaria rudolfii Hoehne
Maxillaria Rufescens Alliance
Maxillaria rufescens Lindl.
Maxillaria rugosa Scheidw.
Maxillaria rugosa Schltr. 1914, not Scheidw. 1843
Maxillaria rupestris Barb.Rodr.

Maxillaria sanderiana Rchb.f.
Maxillaria sanguinea Rolfe
Maxillaria sanguineomaculata Schltr.
Maxillaria sanguinolenta (Lindl.) C.Schweinf.
Maxillaria santanae Carnevali & Ramirez
Maxillaria Saragurensis Alliance
Maxillaria saragurensis Dodson.
Maxillaria saxatilis Rchb.f.
Maxillaria saxicola Schltr.
Maxillaria scabrilinguis Lindl.
Maxillaria Scalariformis Suballiance
Maxillaria scalariformis J.T.Atwood
Maxillaria schistostele Schltr.
Maxillaria schlechteri Foldats
Maxillaria schlechteriana J.T.Atwood
Maxillaria schnitteri Schltr.
Maxillaria schultzei Schltr.
Maxillaria scorpioidea Kraenzl.
Maxillaria scurrilis Rolfe
Maxillaria seidelii Pabst
Maxillaria semiorbicularis Ames & C.Schweinf.
Maxillaria serotina Hoehne, not Regnell & Barb.Rodr.
Maxillaria serotina Regnell & Barb.Rodr.
Maxillaria serrulata Ames & Correll
Maxillaria sessilis (Sw.) Fawc. & Rendle
Maxillaria sessilis Lindl.
Maxillaria setigera Lindl.
Maxillaria seymouriana J.T.Atwood
Maxillaria shepheardii Rolfe
Maxillaria sigmoidea (C.Schweinf.) Ames & Correll
Maxillaria sillarensis Dodson & Vásquez
Maxillaria simacoana Schltr.
Maxillaria similis Garay & Dunsterville
Maxillaria simplicilabia C.Schweinf.
Maxillaria simulans (Ames & C.Schweinf.) L.O.Williams
Maxillaria sinuosa Batem. ex W.Baxt.
Maxillaria skinneri Batem. ex Lindl.
Maxillaria soconuscana D.Breedlove
Maxillaria sodiroi (Schltr.) Dodson
Maxillaria sophronitis (Rchb.f.) Garay
Maxillaria Sophronitis Alliance
Maxillaria sp. nov. ined.
Maxillaria sp. nov., "spotted"
Maxillaria spannagelii Hoehne
Maxillaria spathacea Lindl.
Maxillaria spathulata C.Schweinf.
Maxillaria speciosa Rchb.f.
Maxillaria Speciosa Suballiance
Maxillaria spegazziniana Kraenzl.
Maxillaria spilotantha Rchb.f.
Maxillaria spiritu-sanctensis Pabst
Maxillaria splendens Poepp. & Endl.
Maxillaria Splendens Suballiance
Maxillaria squalens Hook.
Maxillaria squamata Barb.Rodr.
Maxillaria squarrosa (Schltr.) comb. ined.
Maxillaria stachyobiora Rchb.f.
Maxillaria stapelioides (Link & Otto) Lindl.
Maxillaria steelii Hook.
Maxillaria stenobulbon Kl.
Maxillaria stenopetala Know. & Westc.
Maxillaria stenophylla Lehm. & Kraenzl. (1899), not Rchb.f. (1854)
Maxillaria stenophylla Rchb.f., not Lehm. & Kraenzl.
Maxillaria stenostele Schltr.
Maxillaria steyermarkii Fold.
Maxillaria stictantha Schltr.
Maxillaria stictissimum Kraenzl.
Maxillaria striata Rolfe
Maxillaria striatella Kraenzl.
Maxillaria strumata (Endres & Rchb.f.) Ames & Correll
Maxillaria suarezorum Dodson
Maxillaria suaveolens Barringer
Maxillaria subpandurata Schltr.
Maxillaria Subulata Alliance
Maxillaria subulata Lindl.
Maxillaria subulifolia Schltr.
Maxillaria sulcata C.Schweinf., illeg.
Maxillaria sulfurea Schltr.
Maxillaria sulfurina Lem.
Maxillaria superba La Llave & Lex.
Maxillaria superflua Rchb.f.
Maxillaria supina Poepp. & Endl.
Maxillaria surinamensis Focke ex Rchb.f.
Maxillaria swartziana C.D.Adams
Maxillaria synsepala J.T.Atwood

Maxillaria taphallae (Rchb.f.) C.Schweinf.
Maxillaria taracuana Schltr.
Maxillaria tarumaensis Hoehne
Maxillaria tenuifolia Lindl.
Maxillaria tenuis C.Schweinf.
Maxillaria tetragona Lindl.
Maxillaria thurstoniorum Dodson
Maxillaria Tigrina Alliance
Maxillaria tigrina C.Schweinf.
Maxillaria tocotana Schltr.
Maxillaria Tonduzii Suballiance
Maxillaria tonduzii (Schltr.) Ames & Correll
Maxillaria tonsoniae Soto Arenas
Maxillaria torifera (Schltr.) comb. ined.
Maxillaria triangularis Lindl.
Maxillaria tricarinata J.T.Atwood
Maxillaria tricolor Kl., Otto, & Dietr.
Maxillaria tricolor Lindl.
Maxillaria tricolor Ruiz & Pav.
Maxillaria trigona C.Schweinf.
Maxillaria Trilobata Suballiance
Maxillaria trilobata Ames & Correll
Maxillaria trilobulata D.E.Benn. & Christenson
Maxillaria triloris Morren
Maxillaria trinitatis Ames
Maxillaria trinitensis Ames ex W.E.Broadway, name only
Maxillaria triphylla Ruiz & Pav.
Maxillaria tristis Schltr.
Maxillaria truncatilabia Schltr.
Maxillaria truxillensis Rchb.f.
Maxillaria tubercularis J.T.Atwood
Maxillaria tuerckheimii Schltr.
Maxillaria turialbae Schltr.
Maxillaria turneri Nichols.
Maxillaria tutae J.T.Atwood
Maxillaria tyrianthina Hort. ex W.Baxt.

Maxillaria ubatubana Hoehne
Maxillaria umbratilis L.O.Williams
Maxillaria Uncata Alliance
Maxillaria uncata Lindl.
Maxillaria undatiflora Ruiz & pav.
Maxillaria undulata Ruiz & Pav.
Maxillaria unguiculata Schltr.
Maxillaria unguilabia Schltr.
Maxillaria unicarinata C.Schweinf.
Maxillaria urbaniana Lehm. & Kraenzl.

Maxillaria vagans Ames & C.Schweinf.
Maxillaria vaginalis Rchb.f.
Maxillaria Valenzuelana Suballiance
Maxillaria valenzuelana (A.Rich) Nash
Maxillaria valenzuelana ssp. angustata J.T.Atwood
Maxillaria valerioi Ames & C.Schweinf.
Maxillaria vandiformis (Schltr.) C.Schweinf.
Maxillaria vanillodora A.Rich & Gal.
Maxillaria Variabilis Alliance
Maxillaria variabilis Batem. ex Lindl.
Maxillaria variabilis Batem. ex Lindl.
Maxillaria variabilis var. unipunctata Lindl.
Maxillaria varicosa Rchb.f.
Maxillaria variegata Ruiz & Pav.
Maxillaria venusta Lind. & Rchb.f.
Maxillaria verecunda Schltr.
Maxillaria vernicosa Barb.Rodr.
Maxillaria verrucifera C.Schweinf.
Maxillaria verrucifera C.Schweinf.)
Maxillaria vestita Schltr.
Maxillaria villonacensis Dodson
Maxillaria villosa (Barb.Rodr.) Cogn.
Maxillaria vinosa (Schltr.) Senghas
Maxillaria violaceopunctata Rchb.f.
Maxillaria virescens (Schltr.) comb. ined.
Maxillaria virginalis Scheidw.
Maxillaria virguncula Rchb.f.
Maxillaria viridis Lindl.
Maxillaria vitelina lindl.
Maxillaria vitelliniflora Barb.Rodr.
Maxillaria Vittariifolia Alliance
Maxillaria vittariifolia L.O.Williams
Maxillaria vulcanica Lehm. & Kraenzl.

Maxillaria wageneri (Rchb.f.) Rchb.f.
Maxillaria warreana Lodd. ex Lindl.
Maxillaria weberbaueri Schltr.
Maxillaria Wercklei Suballiance
Maxillaria wercklei (Schltr.) L.O.Williams
Maxillaria whittenii Dodson
Maxillaria williamsii Dodson
Maxillaria witsenioides Schltr.
Maxillaria woytkowskii C.Schweinf.
Maxillaria wrightii (Schltr.) Ames & Correll
Maxillaria wrightii (Schltr.) Ames & Correll var. imbricata (Schltr.) Ames & Correll

Maxillaria xanthina Lindl.
Maxillaria xantholeuca Schltr.
Maxillaria xantholeuca var. peruviana C.Schweinf.
Maxillaria xanthorhoda Schltr.
Maxillaria xanthosia Barb.Rodr.
Maxillaria xylobiiflora Schltr.

Maxillaria yanganensis Dodson
Maxillaria yauaperyensis Barb.Rodr.
Maxillaria yzabalana S.Watson


Mormolyca aurorae D.E.Benn. & Christenson
Mormolyca fuchsii J.T.Atwood
Mormolyca galeata (Scheidw.) Garay & Wirth
Mormolyca gracilipes (Schltr.) Garay & Wirth
Mormolyca lineolata Fenzl
Mormolyca peruviana C.Schweinf.
Mormolyca polyphylla Garay & Wirth
Mormolyca ringens (Lindl.) Schltr.
Mormolyca schweinfurthiana Garay & Wirth

Neocogniauxia monophylla (Griseb.) Schlechter

Neourbania Alliance
Neourbania adendrobium (Rchb.f.) Fawc. & Rendle
Neourbania nicaraguensis Hamer & Garay

Ornithidium (Aggregata) Alliance
Ornithidium anceps Rchb.f.
Ornithidium album Hook.
Ornithidium alfaroi Ames & C.Schweinf.)
Ornithidium amparoanum (Schltr.) Pabst
Ornithidium aurantiacum Schltr.
Ornithidium aureum Poepp. & Endl.

Ornithidium bicolor Rolfe
Ornithidium biolleyi Schltr.
Ornithidium bolivianum Schltr.
Ornithidium bracteatum Schltr.
Ornithidium breve Schltr.
Ornithidium broadwayi Cogn.

Ornithidium ceriferum (Barb.Rodr.) Barb. Rodr.
Ornithidium chloroleucum Barb. Rodr.
Ornithidium chrysocycnoides Schltr.
Ornithidium coccineum (Jacq.) Salisb.
Ornithidium compactum Schltr.
Ornithidium conduplicatum Ames & C.Schweinf.
Ornithidium confertum Griseb.
Ornithidium cordyline Rchb.f.)
Ornithidium costaricense Schltr.
Ornithidium croceorubens Rchb.f.
Ornithidium cyperifolium Schltr.

Ornithidium densum (Lindl.) Rchb.f.
Ornithidium dichotomum Schltr.)
Ornithidium distichum Lindl.
Ornithidium divaricatum Barb.Rodr.
Ornithidium dolichophyllum Schltr.

Ornithidium flavoviride (Barb.Rodr.) Barb.Rodr.
Ornithidium fragrans Rolfe
Ornithidium fulgens Rchb.f.)

Ornithidium giganteum Lindl.

Ornithidium heterobulbon Kraenzl.
Ornithidium histrionicum Rchb.f.
Ornithidium huancabambae Kraenzl.

Ornithidium inflexum (Lindl.) Rchb.f.

Ornithidium jamesonii Rchb.f.
Ornithidium jenischianum Rchb.f.

Ornithidium lankesteri Ames
Ornithidium lehmannii Kraenzl.
Ornithidium loefgrenii Cogn.
Ornithidium luteorubrum Lehm. & Kraenzl.

Ornithidium mapiriense Kraenzl.
Ornithidium miniatum Lindl.
Ornithidium multicaule (Poepp. & Endl.) Rchb.f.

Ornithidium nanum Rolfe (not Maxillaria nana Hook.)
Ornithidium neglectum Schltr.
Ornithidium niveum Lindl.
Ornithidium nubigenum Rchb.f.

Ornithidium ochraceum Rchb.f.
Ornithidium olivaceum Kraenzl.

Ornithidium paleatum Rchb.f.
Ornithidium pallidiflavum Schltr.
Ornithidium parviflorum (Poepp. & Endl.) Rchb.f.
Ornithidium parvulum Schltr.
Ornithidium pastoense Schltr.
Ornithidium pendulum cogn.
Ornithidium pfitzerianum Lehm. & Kraenzl.
Ornithidium pittieri Ames
Ornithidium pleuranthioides Schltr.
Ornithidium proliferum (Sw.) Fawc. & Rendle)

Ornithidium ramonense Schltr.
Ornithidium rhodoleucum Schltr.
Ornithidium rhomboglossum Schltr.

Ornithidium sanguinolentum Lindl.
Ornithidium schlechterianum C.Schweinf. [nom. illeg.]
Ornithidium semiscabrum Lindl.
Ornithidium serrulatum Lindl.
Ornithidium sigmoideum C.Schweinf.
Ornithidium simulans Ames & C.Schweinf.
Ornithidium sophronitis Rchb.f.
Ornithidium squamatum (Barb. Rodr.) Barb. Rodr.
Ornithidium stenophyllum Schltr.
Ornithidium strumatum Endres & Rchb.f.

Ornithidium taphallae (Rchb.f.) Rchb.f.
Ornithidium tonduzii Schltr.
Ornithidium toriferum Schltr.

Ornithidium vagans Schltr.
Ornithidium vestitum (Sw.) Rchb.f.
Ornithidium virescens Schltr.

Ornithidium weberbauerianum Kraenzl.
Ornithidium wercklei Schltr.
Ornithidium wrightii (Schltr.) C.Schweinf.

Ornithidium xanthinum Rchb.f., ined.


Pityphyllum amesianum Schltr.
Pityphyllum antioquiense Schltr.
Pityphyllum hirtzii Dodson
Pityphyllum lariciis annum (Kraenzl.) Schltr.
Pityphyllum pinoides H.R.Sweet

Pachyphyllum pamplonense Kraenzl.

Pittierella costaricensis Schltr., invalid

Pittierierella calcarata Schltr.

Pleuranthium adendrobium (Rchb.f.) Benth. & Hook.f.

Pleurothallis valenzuelana A.Rich.

Ponera adendrobium Rchb.f.
Ponera affinis Rchb.f.

Pseudomaxillaria Alliance
Pseudomaxillaria chloroleuca (Barb.Rodr.) Hoehne
Pseudomaxillaria parviflora (Poepp. & Endl.) Brieger
Pseudomaxillaria rhomboglossa (Schltr.) Brieger
Pseudomaxillaria vestita (Sw.) Brieger
Pseudomaxillaria virescens (Schltr.) Brieger

Psittacoglossum atratum Llave & Lex.

Scaphyglottis affinis Poepp. & Endl.
Scaphyglottis parviflora Poeppig & Endl.
Scaphyglottis pendula Poepp. & Endl.
Scaphyglottis ruberrima Lindl.
Scaphyglottis taphallae Rchb.f.

Sepalosaccus Alliance
Sepalosaccus horichii (Senghas) Senghas
Sepalosaccus humilis Schltr.
Sepalosaccus strumatum (Endr. & Rchb.f.) Garay

Siagonanthus multicaulis Poepp. & Endl.


Trigonidium Acuminatum Alliance
Trigonidium acuminatum Batem. ex Lindl.
Trigonidium amesianum
Trigonidium amparoanum Schltr.
Trigonidium aurorae D.E.Benn. & Christenson
Trigonidium brachyglossum (Rich. & Gal.) Schltr.
Trigonidium callistele Rchb.f.
Trigonidium christensonii D.E.Benn.
Trigonidium cucullatum Rchb.f.
Trigonidium Egertonianum Alliance
Trigonidium egertonianum Batem. Ex Lindl.
Trigonidium equitans Garay
Trigonidium grande Garay
Trigonidium insigne Rchb.f. ex Benth. & Hook.
Trigonidium Lankesteri Alliance
Trigonidium lankesteri Ames
Trigonidium Latifolium Alliance
Trigonidium latifolium Lindl.
Trigonidium loretoensis Schltr.
Trigonidium macranthum Barb.Rodr.
Trigonidium monophyllum Griesb.
Trigonidium obtusum Lindl.
Trigonidium obtusum Lindl.
Trigonidium pervuvianum Schltr.
Trigonidium ringens Lindl.
Trigonidium riopalenquense Dodson
Trigonidium seemannii Rchb.f.
Trigonidium spathulatum Lind. & Rchb.f.
Trigonidium subrepens Rolfe
Trigonidium tenue Lodd.
Trigonidium turbinatum Rchb.f.

Zygopetalum bicallosum Rhcb.f.

1John T. Atwood, P.O.Box 566 Woodstock VT 05091

2Germán Carnevali, Centro de Investigación Cientifica de Yucatan (CICY), Ex Hacienda Xcumpich, Vieja Carretera a Progreso, A. P. 87, Cordemex, Merida 97310, Yucatan, México

3Calaway H. Dodson, Missouri Botanical Garden, Box 299, St. Louis, MO 63166-2099 (for correspondence use Selby Gardens' address)

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